Montenegrina dofleini dofleini (Wagner, 1928)

Feher, Zoltan & Szekeres, Miklos, 2016, Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae), ZooKeys 599, pp. 1-137 : 15-17

Taxon classification Animalia Stylommatophora Clausiliidae

Montenegrina dofleini dofleini (Wagner, 1928) Fig. 11H, J

Delima (Montenegrina) dofleini Wagner, 1928 in Hesse 1928: 19.

Montenegrina dofleini dofleini - Nordsieck 1974: 153-154, plate 6, figs 31-32. - Zilch 1981: 127. - Nordsieck 2009: 74.

Montenegrina (Montenegrina) kaiseri Brandt, 1961: 4-5, plate 1, fig. 3.

Montenegrina dofleini kaiseri - Nordsieck 1974: 154, plate 6, fig. 34. - Zilch 1981: 127. - Nordsieck 2009: 74.

Diagnosis.

Shell medium, occasionally with whitish tint. Lower whorls smooth, upper ones very densely striate-costate. Neck weakly inflexed, densely striate-costate. Peristome rounded to ovoid. Lamellae superior and spiralis mostly overlap. In front view lamella inferior more or less well emerged, subcolumellaris mostly visible. Lunella dorsolateral, separate from the short basalis. Subclaustralis and sulcalis absent. Anterior plica superior mostly connected to the lunella complex.

Dimensions

(in mm). Hs: 16.6-21.1, Ws: 4.2-5.5 ( Nordsieck 1974).

Type locality.

"Tomoros, 1500 m" = Macedonia, Galičica Mts, Tomoros Summit [ dofleini ]; Macedonia, Galičica Mts, W of Peštani, ca. 1800 m [ kaiseri ].

Type material.

"Tomoros, 1500 m", leg. Doflein, 24.vii.1918, original series [ dofleini ], syntype status doubtful (ZSM 20150453/2); Peštani, E of the Tomoros Summit, 1800 m, leg. Kaiser, viii.1958, holotype [ kaiseri ] (ZMH 2926, ex SMF 163942), paratypes [ kaiseri ] (SMF 163943/4, SMF 201563, ZHM 2278/8, NMBE 9827/2, NHMW-K 44014/2).

Other material.

Macedonia, Ohrid District, Galičica Mts, ca. 4 km N of the junction at the Sveti Naum to Carina road, 1590 m, 40.9934°N, 20.8569°E, leg. ZF, EH, KJ, HS, 16.x.2014 (NHMW 110430/MN/0038); ca. 600 m E of the Bugarska Chuka, 1650 m, 41.0026°N, 20.8534°E, leg. ZF, EH, KJ, HS, 16.x.2014 (NHMW 110430/MN/0039); ca. 500 m E of the Bugarska Chuka, 1680 m, 41.0019°N, 20.8523°E, leg. ZF, EH, KJ, HS, 16.x.2014 (NHMW 110430/MN/0040); ca. 250 m E of the Bugarska Chuka, 1750 m, 41.0033°N, 20.8488°E, leg. ZF, EH, KJ, HS, 16.x.2014 (NHMW 110430/MN/0041); Bugarska Chuka, E side, 1800 m, 41.0037°N, 20.8470°E, leg. ZF, EH, KJ, HS, 16.x.2014 (NHMW 110430/MN/0042); ca. 300 m W of the Bugarska Chuka, 1740 m, 41.0046°N, 20.8432°E, leg. ZF, EH, KJ, HS, 16.x.2014 (NHMW 110430/MN/0043); ca. 200 m N of the Bugarska Chuka, 1800 m, 41.0051°N, 20.8462°E, leg. ZF, EH, KJ, HS, 16.x.2014 (NHMW 110430/MN/0044); N of the Magaro Summit, 1820 m, 40.9444°N, 20.8270°E, leg. ZE, ZF, JG, 29.vi.2015 (HNHM 99567, NHMW 110430/MN/0046); N of the Magaro Summit, 1870 m, 40.9439°N, 20.8283°E, leg. ZF, 29.vi.2015 (HNHM 99568, NHMW 110430/MN/0047); N of the Magaro Summit, 1900 m, 40.9429°N, 20.8252°E, leg. ZF, 29.vi.2015 (HNHM 99569, NHMW 110430/MN/0048); SE slope of the Galičica Mts, rocky forest along the Carina to Sveti Naum road, 1000-1200 m, leg. LP, PS, AS, 14.vii.1972 (HNHM 36717, SMF 227682); Elšani, above the Elešec Camp, 760 m, 41.0345°N, 20.8105°E, leg. ZF, KJ, HS, 17.x.2014 (NHMW 110430/MN/0045); same locality, leg. LP, PS, AS, 15.vii.1972 (HNHM 36737, HNHM 54405, HNHM 67012, SMF 22768); same locality, leg. ZE, ZF, AH, 7.iv.2004 (HNHM 94433).

Distribution.

This taxon is frequent in the alpine region of the Galičica Mountain. We have found it on the eastern slope of the Bugarska Summit above 1600 m, and on the northern slope of the Magaro Summit above 1800 m. Although sporadically there are occurrences at lower altitudes (e.g. in Elšani at 750 m), on the western slopes of the Galičica, below 1600 m, it is replaced by Montenegrina dofleini fagorum Nordsieck, 1974 (Fig. 14).

Remarks.

According to Wagner (see: Hesse 1928), the material he studied consisted of six shells, only one of them in good condition, which had been dead-collected from the Tomoros, 1500 m, on July 24, 1918. In the ZSM collection there are only two shells, one of these relatively well preserved (Fig. 11H). These are apparently original specimens collected by Doflein. Although no material could be found in the MIZPAS collection (Dominika Mierzwa, personal communication), there is no evidence that the ZSM specimens are part of the material that had been studied by Wagner. Therefore, the syntype status of this sample is doubtful.

Although not synonymizing formally, Nordsieck (2009) has already questioned the subspecific status of kaiseri . Two arguments seem to support Nordsieck’s opinion that kaiseri could be regarded as junior synonym of dofleini . First, according to the label information, the type series of kaiseri were collected in the vicinity of the Tomoros Summit, which is the type locality of dofleini . Second, the paratypes of kaiseri show high variation in the traits defined by Brandt as distinguishing features.

In the original description the type locality of kaiseri was erroneously defined as 'W of Peštani’ ( Brandt 1961), because the Galičica range is east of this village.

The type series of kaiseri was collected by Kaiser, the curator of the ZMH Mollusc Collection at that time, and the entire lot was originally housed in the ZMH. As indicated in the description, Brandt (1961) intended depositing the holotype in the SMF. As this would have been done without authorization, the type was returned to the ZMH ( Zilch 1981, Bernhard Hausdorf, personal communication).