Phlegmariurus reflexus (Lam.) B.Øllg., Rodriguésia

Øllgaard, Benjamin, Kessler, Michael & Smith, Alan R., 2018, Prodromus of a fern flora for Bolivia. II. Lycopodiaceae, Phytotaxa 334 (3), pp. 255-294 : 283

Phlegmariurus reflexus (Lam.) B.Øllg., Rodriguésia 63 481. 2012. ( Fig. 14C )

= Lycopodium reflexum Lam., Encycl. 3: 653. 1792.

= Huperzia reflexa (Lam.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248. 1874.

= Lycopodium pearcei Baker, Handb. Fern-Allies 14. 1887.

= Huperzia pearcei (Baker) Holub, Folia Geobot. Phytotax. 20: 75. 1985.

= Phlegmariurus pearcei (Baker) B.Øllg., Phytotaxa 57: 17. 2012.

Range: — Phlegmariurus reflexus is widespread throughout the Neotropics, and occurs from southern Mexico south to Bolivia (BE, CH, CO, LP, SC) and Argentina.

Ecology: —Common; frequent pioneer on permanently moist, disturbed ground, e.g., road banks, but also in peat bogs, in humid to semideciduous forests; 500–3000 m, 300–3400 m elsewhere.

Notes: — Phlegmariurus reflexus is highly variable with respect to leaf and stem size, direction and crowding of leaves, and leaf margin characters. Part of the variation may reflect variable growth conditions, but very often several distinct forms may co-occur in the same habitat, indicating that genetic differences exist. The variation patterns are complex and in need of a detailed study.

A small form was recognized earlier as var. minor , but appears to intergrade with the remaining species complex, and therefore is abandoned at this point.

Phlegmariurus pearcei was included as a separate species in Øllgaard & Gonzales (2014: 126) but is included in P. reflexus here.

Figures

FIGURE 14. A, Phlegmariurus eversus, growth habit, A1, close-up of sporangiate division (Holm-Nielsen 5644, Ecuador, AAU), A2, close-up of sporangiate division (Holmgren 674, Ecuador, S).—B, Phlegmariurus hypogaeus, growth habit, B1, close-up of sporangiate division (Holm-Nielsen et al. 5343, Ecuador, AAU).—C, Phlegmariurus reflexus, growth habit, C1, close-up of sporangiate division (Holm-Nielsen et al. 3873, Ecuador, AAU), C2, close-up of sporangiate division (Holm-Nielsen et al. 207, Ecuador, AAU).