Canoptila williami, Robertson, Desiree R. & Holzenthal, Ralph W., 2006

Robertson, Desiree R. & Holzenthal, Ralph W., 2006, The Neotropical caddisfly genus Canoptila (Trichoptera: Glossosomatidae), Zootaxa 1272, pp. 45-59 : 53-55

Canoptila williami , new species

(Figs. 4,5)

Abdominal segment IX in Canoptila williami is narrower laterally and ventrally than in C. bifida . Tergum X in lateral view is quadrate in C. williami and triangular in C. bifida . In the new species tergum X is subquadrate in dorsal view while it is bifid in C. bifida . On the endotheca, C. williami has larger parameres, and smaller, less sclerotized dorsomesal processes than C. bifida . In the forewing of C. williami , the apical part of Cu2 turns sharply downward to meet the wing margin apicad of the intersection of Cu1 and Cu2, instead of at the intersection as in C. bifida .

Adult. Length of forewing: male 2.5–3.0 mm (n =5); female 2.6–3.2 mm (n =4). Body, wings, and appendages nearly uniformly fuscous, tibia and tarsi yellowish brown. Wings ( Figs. 2 A, 2B) as detailed in the generic diagnosis, except apical part of Cu2, turning sharply down to meet the wing margin apicad of the intersection of Cu1 and Cu2.

Male genitalia ( Fig. 4 ). Sternum VI ( Fig. 4 A) with thin, digitate mesal process, projecting caudally. Abdominal segment IX ( Fig. 4 B) well developed dorsally, narrow laterally and ventrally, anterior margin nearly straight. Tergum IX in lateral view ( Fig. 4 B), with anterior margin straight, forming truncate ridge apicodorsally; in dorsal view ( Fig. 4 C), posterior margin subtriangular, covered in fine microtrichia. Tergum X in lateral view ( Fig. 4 B), quadrate, posterior and ventral margins irregular, setose dorsally, apically, and ventrally, apex subtruncate; in dorsal view ( Fig. 4 C), subquatrate, slightly irregular, with pair of long, straight, spine­like processes arising from posterolateral margin and forming an apparent articulation, directed inward. Phallocrypt ( Fig. 4 B) extending from beneath tergum X to ventral region of segment IX, entirely membranous except for lightly sclerotized phallic shield where phallic apparatus is attached. Phallobase ( Figs. 4 B, 4D) reduced, lightly sclerotized, with paired patches of setae ventrolaterally. Endotheca ( Figs. 4 B, 4D) membranous, basoventrally bearing pair of large, membranous digitate parameres with large globular, asperous apices. Endophallus ( Figs. 4 B, 4D) highly membranous, enlarged and convoluted when evaginated; apicomesally, bearing single pair of sharply pointed, sclerotized processes; apicodorsally, bifid, with membranous upper lobe, lightly sclerototized lower lobe.

Female genitalia ( Figs. 5 A & 5B). Sternum VI with thin digitate mesal process, projecting caudally. Tergum VI and VII with patches of long, bristle­like setae posteriorly. Abdominal segment VIII synscleritous, relatively short. Segments IX and X closely associated to the point that their boundaries cannot be discerned with any certainty, with pair of small digitate cerci dorsolaterally. Genital chamber lightly sclerotized, short, extending to anterior end of segment VIII, with pair of membranous lobes ventrally. Vaginal apparatus sclerotized; in lateral view ( Fig. 5 A), suboval, with slender process at posterior end extending into genital chamber; in ventral view ( Fig. 5 B), convex and keelshaped, with rectangular posterior end and subtriangular anterior end.

Figures

FIGURE 2. Canoptila bifida Mosely—holotype (UMSP 000107591). Wings: A. forewing; B. hind wing.

FIGURE 4. Canoptila williami n. sp. (UMSP 000087912). Male: A. process of sternum VI; B. genitalia, lateral; C. genitalia, dorsal; D. genitalia, ventral.

FIGURE 5. Canoptila williami n. sp. (UMSP 000029632). Female genitalia: A. lateral, B. ventral.