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Centruroides Marx, 1890

Goodman, Aaron M., Prendini, Lorenzo, Francke, Oscar F. & Esposito, Lauren A., 2021, Systematic Revision Of The Arboreal Neotropical “ Thorellii ” Clade Of Centruroides Marx, 1890, Bark Scorpions (Buthidae C. L. Koch, 1837) With Descriptions Of Six New Species, Bulletin of the American Museum of Natural History 2021 (452), pp. 1-93 : 6-8

publication ID

 https://doi.org/ 10.1206/0003-0090.452.1.1

DOI

 https://doi.org/10.5281/zenodo.5825900

persistent identifier

https://treatment.plazi.org/id/510E87A5-6D54-FF99-2410-FD71FB05FBC7

treatment provided by

Carolina

scientific name

 Centruroides Marx, 1890
status

 

Centruroides Marx, 1890 View in CoL

Figures 1–43 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 View FIGURE 12 View FIGURE 13 View FIGURE 14 View FIGURE 15 View FIGURE 16 View FIGURE 17 View FIGURE 18 View FIGURE 19 View FIGURE 20 View FIGURE 21 View FIGURE 22 View FIGURE 23 View FIGURE 24 View FIGURE 25 View FIGURE 26 View FIGURE 27 View FIGURE 28 View FIGURE 29 View FIGURE 30 View FIGURE 31 View FIGURE 32 View FIGURE 33 View FIGURE 35 View FIGURE 37 View FIGURE 38 View FIGURE 39 View FIGURE 41 View FIGURE 42 View FIGURE 43 ; tables 1–10 View TABLE 1

Centrurus (nec Ehrenberg, 1829) : Thorell, 1876a: 9; 1876b: 83; Karsch, 1879a: 18; Pocock, 1890: 120, 121, 127; Kraepelin, 1891: 119– 124 (part); Pocock, 1893: 375, 385, 386; Laurie, 1896: 131; Lönnberg, 1897: 196, 197, 208; Kraepelin, 1899: 87 (part); Banks, 1900: 425; Borelli, 1909: 222; Comstock, 1912: 25, 27, fig. 31; Birula, 1917a: 164; 1917b: 54, 107; Ochoterena, 1920: 223; Mello-Campos, 1924a: 246; 1924b: 312; Comstock, 1940: 27, fig. 31 (lapsus calami); Millot and Vachon, 1949: 427; Díaz Nájera, 1966: 110, 111, pl. 1; 1970: 113.

Centruro: Karsch, 1879b: 120 (lapsus calami).

Centruroides Marx, 1890: 211 View in CoL , type species Buthus exilicauda Wood, 1863 (= Centruroides exilicauda ( Wood, 1863) , by monotypy; Pocock, 1902a: 19, 20; 1902b: 365; Kraepelin, 1912: 69–71; 1914: 22; Hoffmann, 1932: 244, 245; Mello-Leitão, 1932: 27; 1934: 4, 5; Franganillo, 1936: 158; Hoffmann, 1937: 201–203; Moreno, 1939: 63; Comstock, 1940: 25, 27; Moreno, 1940: 164; Mello-Leitão, 1942: 126; 1945: 240, 250–252; Scorza, 1954:190; Bücherl, 1964: 59; 1967: 113; Muma, 1967: 2, 4; Bücherl, 1969: 767; Aguilar and Meneses, 1970: 3; Bücherl, 1971: 327; Stahnke, 1971: 282; 1972: 125, fig. 9; Armas, 1974a: 25; Vachon, 1974: 906, 908; 1975: 1598; Francke, 1977: 75; Stahnke and Calos, 1977: 111; Vachon, 1977: 294; Lourenço, 1979: 214; Williams, 1980: 2, 4; Armas, 1984: 2; González-Sponga, 1984: 64, 65; Francke, 1985: 7, 15; Francke and Stockwell, 1987: 7; Armas, 1988: 44, 91, 95; Stockwell, 1988: 3; Sissom, 1990: 94, 101; Nenilin and Fet, 1992: 9, 12–14; Stockwell, 1992: 412, 419; González-Sponga, 1996: 118, 119, 124, 125, figs. 285–287, 289, 292; Armas, 1998: 50; Kovařík, 1998: 106; Fet and Lowe, 2000: 98–122; Towler et al., 2001: 161–163; Armas et al., 2004: 167–171, tables 1–3 View TABLE 1 View TABLE 2 View TABLE 3 ; Prendini and Wheeler, 2005: 481, table 10 View TABLE 10 ; Kamenz and Prendini, 2008: 6, 8, 22, 40, tables 1 View TABLE 1 , 2 View TABLE 2 , pl. 13–17; Volschenk et al., 2008: 654, 656, 658, 659, 663, 664, 674, fig. 1C, tables 1 View TABLE 1 , 2 View TABLE 2 ; Armas et al., 2012: 106, 112; Cupitra Vergara et al., 2014: 207–215, figs. 1–4, tables 1 View TABLE 1 , 2 View TABLE 2 ; Kovařík and Teruel, 2014: 1, 7, 15, 25, 26; Loria and Prendini, 2014: 3, 9, 10, 24, 25, fig. 2D, table 5 View TABLE 5 ; Lourenço, 2014: 60, 63; Miller et al., 2014: 301, 306, 307, table 1 View TABLE 1 ; Ponce-Saavedra and Francke, 2014: 54, figs. 11, 14; Quintero Arias and Esposito, 2014: 373–382, figs. 1–6, table 1 View TABLE 1 ; Teruel and Rodríguez-Cabrera, 2014: 131; Armas and Ávila, 2015: 66, 67, 69, 70, 71; Ponce-Saavedra et al., 2015: 81–89, figs. 1–5, tables 1 View TABLE 1 , 2 View TABLE 2 ; Quijano-Ravell and Ponce-Saavedra, 2015: 35–44, figs. 1–5, tables 1–3 View TABLE 1 View TABLE 2 View TABLE 3 ; Riquelme et al., 2015: 2, 16, 18, 19; Teruel et al., 2015a: 3–14, figs. 1–37, table 1 View TABLE 1 ; 2015b: 1–18, figs. 1–35, tables 1–5 View TABLE 1 View TABLE 2 View TABLE 3 View TABLE 4 View TABLE 5 ; 2015c: 13–33, figs. 1–75, tables 1–4 View TABLE 1 View TABLE 2 View TABLE 3 View TABLE 4 ; Baldazo-Monsivaiz et al., 2016: 76–79, fig. 2A, table 1 View TABLE 1 ; de los Santos et al., 2016: 1–7, 14, 16–19, 22, figs. 2A–G, 3A, table 1 View TABLE 1 ; Kovařík et al., 2016a: 1, 11, 18, 19; Lourenço and Velten, 2016: 61, 65; Quijano-Ravell and Ponce-Saavedra, 2016a: 49–61, figs. 1–9, tables 1–4 View TABLE 1 View TABLE 2 View TABLE 3 View TABLE 4 ; 2016b: 85–90, figs. 1–4; Teruel, 2016: 91–93, figs. 1–3; Baldazo-Monsivaiz, 2017: 21–27, fig. 2A; Esposito et al., 2017: 2–15 View Cited Treatment , 29, 30, 35–37, 41, 61, 71, 81, 85, 101, 116, 122, 124, 126, 128–130, fig. 13; McWest et al., 2015: 3, 9, 10, 25, 28, 30–34, 38, figs. 2A, 3A, 4A; Teruel and Myers, 2017: 1–14, figs. 1–11, table 1 View TABLE 1 ; Teruel et al., 2017: 6; Ubinski et al., 2018: 475–486, figs. 1–8, tables 1 View TABLE 1 , 2 View TABLE 2 ; Esposito and Prendini, 2019: 1–10, figs. 1, 2, 3, tables 1–4 View TABLE 1 View TABLE 2 View TABLE 3 View TABLE 4 ; Crews and Esposito, 2020: 8, 9, 12, 17, 18; Fet and Kovařík, 2020: 8, 10, 12, 14, 18, 19, 22–26, 28, 29, 31; Howard et al., 2019: 74, 84, fig. 3, table 1 View TABLE 1 ; Miranda and Armas, 2020: 5, 7; Ponce-Saavedra and Francke, 2019: 1–17, figs. 1–11, tables 1–4 View TABLE 1 View TABLE 2 View TABLE 3 View TABLE 4 ; Quijano-Ravell et al., 2019: 31–48, figs. 1–32, tables 1–5 View TABLE 1 View TABLE 2 View TABLE 3 View TABLE 4 View TABLE 5 ; Teruel and Questel, 2020: 12, 14; Trujillo and Armas, 2020: 1–8, figs. 1–4, table 1 View TABLE 1 ; Wendruff et al., 2020: 2, 5, 6, fig. 2B.

Centruroides (Centruroides) : Werner, 1934: 273.

Centruoroides: Díaz Nájera, 1970: 117 (lapsus calami).

Centruriodes: Lourenço and Eickstedt, 1988: 7 (lapsus calami).

Key to Identification of the Species of the “ thorellii ” Clade of Centruroides Marx, 1890

1. Carapace posterosubmedian carinae............2

– Carapace posterosubmedian carinae absent...6

2. Pedipalp chela manus, retrodorsal carina smooth, dorsomedian carina weakly granular; mesosomal sternite VII, ventrolateral and ventrosubmedian carinae obsolete to absent; metasomal segments I–III, ventrosubmedian and ventrolateral carinae vestigial, smooth (figs. 18, 19D); telson vesicle surfaces smooth (♀) (figs. 23–25D)............. .................... Centruroides rileyi Sissom, 1995

– Pedipalp chela manus, retrodorsal carinae present and dorsomedian carinae present or restricted to proximal third or half; mesosomal sternite VII, ventrolateral and ventrosubmedian carinae distinct; metasomal segments I–III, ventrolateral and ventrosubmedian carinae complete, granular (figs. 18L, R, 19L, R); telson vesicle surfaces granular (♀) (figs. 23L, R, 24L, R, 25L, R).....................3

3. Pedipalp chela manus, prodorsal carina absent ....................................... 4

– Pedipalp chela manus, prodorsal carina present .................................... 5

4. Pectinal tooth counts, 15–17 (♂), 14 (♀) ( table 9 View TABLE 9 ); mesosomal tergites I–VII, dorsomedian carinae present on I–III, absent on IV–VII; mesosomal sternite VII, ventrolateral and ventrosubmedian carinae distinct; metasoma length greater than 3× mesosoma length; metasomal segments I–IV, ventrolateral and ventrosubmedian carinae finely granular on segments I–III (figs. 18I, L, 19I, L), smooth on IV....... .................... Centruroides chanae , sp. nov.

– Pectinal tooth counts, 14 (♂), 12–14 (♀) ( table 10 View TABLE 10 ); mesosomal tergites I–VII, dorsomedian carinae absent on I and II, vestigial on III–VI, absent on VII; mesosomal sternite VII, ventrolateral and ventrosubmedian carinae weakly developed; metasoma length less than 3× mesosoma length; metasomal segments I–IV, ventrolateral and ventrosubmedian carinae present on segments I–IV (figs. 18O, R, 19O, R)....... Centruroides yucatanensis , sp. nov.

5. Mesosomal tergites I–VII, dorsomedian carinae distinct, complete; total length (♂, ♀), 44–60 mm ( table 8 View TABLE 8 ); pectinal tooth counts, 13–15 (♂), 13 (♀)............................... ... Centruroides hoffmanni Armas, 1996

– Mesosomal tergites I–VII, dorsomedian carinae restricted to posterior two-thirds of segments I–VI, absent on VII; total length (♂, ♀), 38–52 mm ( table 7 View TABLE 7 ); pectinal tooth counts, 14–16 (♂), 12–14 (♀)....................... .............. Centruroides schmidti Sissom, 1995

6. Pedipalp chela manus, prodorsal carina complete, granular...................................... ....... Centruroides cuauhmapan , sp. nov.

– Pedipalp chela manus, prodorsal carina absent ........................................ 7

7. Pedipalp chela manus, retrodorsal carina complete, granular; mesosomal sternite VII, ventrolateral and ventrosubmedian carinae obsolete to absent (figs. 13, 14A); telson vesicle surfaces (♀) smooth (figs. 23–25E)...... .................. Centruroides hamadryas , sp. nov.

– Pedipalp chela manus, retrodorsal carina weakly granular; mesosomal sternite VII, ventrolateral and ventrosubmedian carinae weakly developed; telson vesicle surfaces (♀) granular.......8

8. Total length, 39–51 mm ( table 6 View TABLE 6 ); pedipalp chela manus, dorsomedian carina weakly granular, restricted to distal half; pectinal tooth counts, 13–14 (♂, ♀); mesosomal tergites I–VII, dorsomedian carinae vestigial; metasoma length less than 3.2× greater than mesosoma length; metasomal segments I– IV, ventrolateral and ventrosubmedian carinae absent (figs. 18M, P, 19M, P)................ ............. Centruroides catemacoensis , sp. nov.

– Total length, 48–55 mm ( table 5 View TABLE 5 ); pedipalp chela manus, dorsomedian carina absent; pectinal tooth counts, 14–16 (♂), 13–15 (♀); mesosomal tergites I–VII, dorsomedian carinae weakly granular, restricted to posterior half; metasoma length greater than 3.2× mesosoma length; metasomal segments I– IV, ventrolateral and ventrosubmedian carinae present (figs. 18H, K, 19H, K).......... ........................ Centruroides berstoni , sp. nov.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Cerambycidae

Loc

Centruroides Marx, 1890

Goodman, Aaron M., Prendini, Lorenzo, Francke, Oscar F. & Esposito, Lauren A. 2021
2021
Loc

Centruriodes: Lourenço and Eickstedt, 1988: 7

Lourenco, W. R. & V. R. D. von Eickstedt 1988: 7
1988
Loc

Centruoroides: Díaz Nájera, 1970: 117

Diaz Najera, A. 1970: 117
1970
Loc

Centruroides Marx, 1890: 211

Crews, S. C. & L. A. Esposito 2020: 8
Fet, V. & F. Kovarik 2020: 8
Miranda, R. J. & de Armas, L. F. 2020: 5
Esposito, L. A. & L. Prendini 2019: 1
Howard, R. J. & G. D. Edgecombe & D. A. Legg & D. Pisani & J. Lozano-Fernandez 2019: 74
Ponce-Saavedra, J. & O. F. Francke 2019: 1
Baldazo-Monsivaiz, J. G. & R. Teruel & I. Canche-Aguilar 2017: 21
Esposito, L. A. & H. Y. Yamaguti & C. A. Souza & R. Pinto-Da- & L. Prendini 2017: 2
Baldazo-Monsivaiz, J. G. 2016: 76
Santos, G. & L. F. de Armas & R. Teruel 2016: 1
Kovarik, F. & G. Lowe & J. Pliskova & F. Stahlavsky 2016: 1
Lourenco, W. R. & J. Velten 2016: 61
Armas, L. F. & A. F. Avila 2015: 66
McWest, K. J. & Z. J. Valois & W. D. Sissom 2015: 3
Cupitra Vergara & N. I & S. Cubides Cubillos & M. M. Saldarriaga-Cordoba & E. Estrada-Gomez 2014: 207
Kovarik, F. & R. Teruel 2014: 1
Loria, S. F. & L. Prendini 2014: 3
Lourenco, W. R. 2014: 60
Miller, A. L. & R. A & Makowsky, D. R. & Formanowicz, L. & C. L. Cox 2014: 301
Ponce-Saavedra J. & O. F. Francke 2014: 54
Armas, L. F. de & D. Luna-Sarmiento & E. Florez 2012: 106
Kamenz, C. & L. Prendini 2008: 6
Armas, L. F. de & E. Martin-Frias & J. F. Paniagua-Solis 2004: 167
Fet, V. & G. Lowe 2000: 98
Armas, L. F. de 1998: 50
Kovarik, F. 1998: 106
Gonzalez-Sponga, M. A. 1996: 118
Nenilin, A. B. & V. Fet 1992: 9
Armas, L. F. de 1988: 44
Francke, O. F. & S. A. Stockwell 1987: 7
Francke, O. F. 1985: 7
Armas, L. F. de 1984: 2
Gonzalez-Sponga, M. A. 1984: 64
Lourenco, W. R. 1979: 214
Francke, O. F. 1977: 75
Armas, L. F. de 1974: 25
Bucherl, W. 1971: 327
Aguilar, F. P. G. & G. O. Meneses 1970: 3
Bucherl, W. 1969: 767
Bucherl, W. 1967: 113
Muma, M. H. 1967: 2
Bucherl, W. 1964: 59
Mello-Leitao, C. F. de 1945: 240
Mello-Leitao, C. F. de 1942: 126
Comstock, J. H. 1940: 25
Moreno, A. 1940: 164
Moreno, A. 1939: 63
Hoffmann, C. C. 1937: 201
Franganillo, P. B. 1936: 158
Mello-Leitao, C. de 1934: 4
Hoffmann, C. C. 1932: 244
Mello-Leitao, C. F. de 1932: 27
Kraepelin, K. 1912: 69
Pocock, R. I. 1902: 19
Marx, G. 1890: 211
1902b: 365
1914: 22
Scorza, 1954:190
Stahnke, 1971: 282
1972: 125 , fig. 9
Vachon, 1974: 906 , 908; 1975: 1598
Stahnke and Calos, 1977: 111
Vachon, 1977: 294
Williams, 1980: 2 , 4
Stockwell, 1988: 3
Sissom, 1990: 94 , 101
Stockwell, 1992: 412 , 419
1890
Loc

Centruro:

Karsch, F. 1879: 120
1879
Loc

Centrurus (nec Ehrenberg, 1829)

Diaz Najera, A. 1966: 110
Millot, J. & M. Vachon 1949: 427
Comstock, J. H. 1940: 27
Mello-Campos, O. de 1924: 246
Mello-Campos, O. de 1924: 312
Ochoterena, I. 1920: 223
Birula, A. A. 1917: 164
Birula, A. A. 1917: 54
Comstock, J. H. 1912: 25
Borelli, A. 1909: 222
Banks, N. 1900: 425
Kraepelin, K. 1899: 87
Lonnberg, E. 1897: 196
Laurie, M. 1896: 131
Pocock, R. I. 1893: 375
Kraepelin, K. 1891: 119
Pocock, R. I. 1890: 120
Karsch, F. 1879: 18
Thorell, 1876a: 9
1876b: 83
1970: 113

Specimens

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