Eunatalis Schenkling, 1909

Gerstmeier, Roland & Seitner, Martin, 2013, Revision of the checkered beetle genus Eunatalis Schenkling, 1909 (Coleoptera: Cleridae: Clerinae), Zootaxa 3698 (1), pp. 1-77 : 6

Eunatalis Schenkling, 1909

Type species: Notoxus porcatus Fabricius, 1787: 127 .

Schenkling 1909: 163; Elston 1921: 148; Kolibáč 1998: 191; Solervicens 2007: 57.

Head: Dark brown to black; vested with whitish to grey setae; labrum broadly emarginate, V-shaped, light brown to dark brown; terminal maxillary palpomeres with sides parallel to very weakly divergent and apex obliquely truncate, terminal labial palpomeres securiform ( Fig. 1 ); eyes coarsely facetted, protruding laterally, margined; head including eyes as broad as or broader than anterior width of pronotum; interocular space from one to double eye width (as viewed from above); gular sutures converging, gular process conspicuous, with two papilles. Antennae with 11 antennomeres, filiform, light brown to black; antennomeres with dense, fine punctation, terminal three antennomeres densely tomentose, forming a loose club, A 11 in E. spinicornis acuminate ( Fig. 2 ).

Thorax: Light brown to black; punctation and pilosity variable, base and apex with fringe of setae, pilosity mostly grey to whitish; pronotum longer than wide, sides more or less sinuate, with wrinkles, middle always with an elongated groove (sometimes missing or weak); grooves of sub-basal collar on each side of pronotum fused or separated by a weak or distinct bar. Pro-intercoxal process narrow, broadly dilated distally; procoxal cavities opened to almost closed posteriorly (sometimes variable within a species); hypomeron tapering apically towards intercoxal process; mesoventrite with variable sculpture and punctation, anterior mesoventral process distinct, sinuate, pit-shaped, slightly opened apically; posterior part of mesepisternon mostly smooth or slightly punctate ( Fig. 3 ); metaventrite with a conspicuous discriminal line, more or less punctate and wrinkled. Scutellum round to oblong-ovate, light brown to black, with isolated lateral setae, slightly punctate. Metendosternite with conspicuous furcal arms, laminae broadly triangular ( Fig. 4 ).

Elytra: Light brown to black; pilosity variable, mostly densely vested with anteriorly directed setae at the lower margin of elytrae, whitish to grey; elytral base not margined, lateral margins parallel or broadest behind middle, apices broadly rounded or with a conspicuous tooth, elytral punctation arranged into ten striae ( E. laevis with hidden punctures), the ninth stria is partly or completely vestigial, interstices with a fine microsculpture (wrinkled to punctate); diameter of punctations from base to apex similar, or diameter of punctations behind the middle decreasing or obsolete; punctation round or oblong, with two or three nodules (a pair of lateral nodules, and one posterior nodule), nodules more or less sinuate, with a single seta beneath the basal nodule ( Fig. 5 ); interstices in basal half flattened, posterior of middle sometimes more or less ridged. Hindwing with closed wedge cell, margin of radial cell strongly sclerotized, MP3, MP4, AA3+4 and CuA+AA reaching margin ( Fig. 7 ).

Legs: Light brown to black; more or less pilose, inner edge of protibiae and sometimes mesotibiae with a row of short, thick bristles, with more or less conspicuous punctation; profemora conspicuously swollen, protibiae slightly bent, mesotibiae more or less bent (except E. semicostata with almost straight mesotibiae), metatibiae straight to very slightly bent, all tibiae without longitudinal carinae, protibiae terminating in a short spine and a single spur, tibial spur formula 1-2-2; tarsal pulvillar formula 3-3-3, tarsal pulvilli broadly lobed, but not emarginate; claws simple ( Fig. 6 ).

Abdomen: Six visible abdominal ventrites, with a more or less dense pilosity and punctation. Ventrite VI in males straight or emarginate, in females rounded. Tegmen ventrally closed, phallobasic apodeme and struts distally more or less spoon-like, branches of spicular fork distally strongly broadened. Female copulatory organs have not been investigated.

Figures

FIGURES 1 – 10. Morphological characters of the genus Eunatalis. (1) Head, lower surface (Scale bar 1 mm), (2) antennae (Scale bar 1 mm), (3) Pro- and mesothorax, lower surface (Scale bar 1 mm), (4) Metendosternite (Scale bar 1 mm), (5) Nodules of elytral punctation, arrow indicates the basal seta (Scale bar 20 µm), (6) Protarsus, with spine, spur and pulvilli (Scale bar 1 mm), (7) Wing venation (Scale bar 5 mm), (8) Longitudinal groove between eyes of E. ninae n. sp. (Scale bar 1 mm), (9) Elytral punctation with two lateral nodules, E. foveata (Scale bar 1 mm), (10) Elytral punctation with two lateral nodules and one basal nodule, E. pernodulosa n. sp. (Scale bar 1 mm).