Argemiones stupeflip, BRAZIDEC, 2023

BRAZIDEC, MANUEL, 2023, Argemiones stupeflip gen. et sp. nov., a new spathiopterygid wasp (Hymenoptera: Diaprioidea) from the mid-Cretaceous Kachin amber, Palaeoentomology 6 (3), pp. 292-299 : 294-297

Argemiones stupeflip sp. nov.

( Figs 2–4 ) urn:lsid:zoobank.org:act:BA3F9D4C-8803-4BF8-AE49-05C75129137C

Material. Holotype male IGR.BU-076, a well-preserved male, only lacking left pedicel, flagellomeres 1 and 2; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR).

Etymology. This species is dedicated to the French musical band Stupeflip, exploring various genres such as hip-hop, punk, electronic music and pop. The specific epithet is to be treated as a name in apposition.

Diagnosis. As for genus.

Type locality and horizon. Noije Bum , Hukawng Valley , Kachin State, northern Myanmar; late Albian-early Cenomanian, ca. 99 Ma, mid-Cretaceous .

Description. Body length 0.76 mm; integument dark castaneous.

Head hypognathous, globular (length 0.16 mm, width above eyes 0.23 mm); back of head coarsely punctured; compound eye bulging, occupying about half of head lateral surface, composed of numerous ommatidia (number over 60 for each eye); torulus inserted on frontal shelf, higher than eye; antenna filiform, longer than body; scape elongate, longer than F1 (0.10 mm); pedicel globular, about twice as long as wide (length 0.05 mm); 12 flagellomeres cylindrical, elongate, F1 and F2 not modified, flagellomeres 1–11 progressively shortening (0.10 mm> F1> 0.09 mm> F2–F4> 0.08 mm> F5–F6> 0.07 mm> F7–F10> 0.06 mm> F11) and widening, flagellomere 12 acuminate, about as long as flagellomere 4; pubescence on flagellomeres denser toward apex of antenna, setae fine, not longer than a third of flagellomere length.

Mesosoma stout (length 0.29 mm); mesoscutum large, convex, about half length of mesosoma, apparently smooth; notaulus present, convergent posteriorly; pronotum apparently without lateral striations, not visible in dorsal view.

Fore wing paddle-like, longer than body (length 1.13 mm); membrane slightly wrinkled medially, smooth apically, uniformly micropubescent; fringe of long setae (longest 0.07 mm) present along apical margin; parastigma sclerotized and slightly thickened, extending approximately to half of wing; R, Rs, M, Cu and A veins spectral but visible apically, outlined by microtrichiae, not sclerotized basally; fork of Rs and M basad apex of parastigma; A longer than parastigma; B not sclerotized. Hind wing strongly reduced, only present as remnant of vein.

Legs slender, long with respect to body, with sparse setae; tibia with inner row of setae; tibial spur formula 1-1-1, metatibial spur longest and slightly curved; probasitarsomere shorter than remainder of protarsus; meso- and metabasitarsomeres, respectively, distinctly longer than remainder of meso- and metatarsus; tarsal claws simple, angled apically; arolium well-developed.

Petiole visible, not obscured by gaster, slightly longer than wide (length 0.06 mm, width 0.05 mm), smooth.

Gaster globose (length 0.25 mm); second metasomal tergite and sternite longest, hiding other tergites and sternites.

Key to the genera and species of Spathiopterygidae

2. Compound eye bulging with few ommatidia (‘berry-like’), parastigma thin (thinner than other visible veins)..................................3 [ Spathiopteryx Engel & Ortega-Blanco, 2013 ]

- Compound eye with numerous ommatidia (more than 50 ommatidia), parastigma thick (at least as thick as other visible veins).......................................................................4

3. Compound eye with about 40 ommatidia, pronotum with transverse striation, vertex punctured........... Spathiopteryx alavarommopsis Engel & Ortega-Blanco, 2013

- Compound eye with about 20 ommatidia, pronotum smooth, vertex punctate to areolate.............................. Spathiopteryx soosi Szabó, Brazidec & Perrichot, 2022

4. Fore wing membrane entirely wrinkled, with a positive gradient of microtrichiae density towards apex.................. .......... Diameneura marveni Santer & Álvarez-Parra, 2022

- Fore wing membrane partly smooth, microtrichiae distributed evenly on membrane....................................5

5. Fore wing with M+Cu desclerotized, hind wing reduced as a venal stub ............. Argemiones stupeflip gen. et sp. nov.

- Fore wing with M+Cu sclerotized, hind wing with membrane and hamuli 6 [ Mymaropsis Engel & Ortega-Blanco, 2013 ]

6. Scape slightly expanded ventrally, pedicel globular, flagellomeres 5–12 twice as wide as flagellomeres 1– 4 ....................................................................... Mymaropsis baabdensis Krogmann, Azar, Rajaei & Nel, 2016

- Scape not ventrally expanded, pedicel almost globular, flagellomeres evenly wide.................................................. ....... Mymaropsis turolensis Engel & Ortega-Blanco, 2013

7. Head coarsely punctured, antennae short, not longer than body length, gaster globose, nearly as long as mesosoma ........................................................ Spathiopria sayrevillensis Engel, Ortega-Blanco & Grimaldi, 2013

- Head imbricate, antennae elongate, longer than body length, gaster ovoid, longer than mesosoma........................... ......................... Diaspathion ortegai Engel & Huang, 2015

Figures

FIGURE 2. Argemiones stupeflip gen. et sp. nov., specimen IGR.BU-076, habitus in dorsal view. Scale bar = 1 mm.

FIGURE 3. Argemiones stupeflip gen. et sp. nov., specimen IGR.BU-076, line drawing of habitus in dorsal view (pst = parastigma). Scale bar = 1 mm.

FIGURE 4. Argemiones stupeflip gen. et sp. nov., specimen IGR.BU-076, A, Habitus in ventral view. B, Right fore wing and metasoma in dorsal view (arrow = remnant of hind wing). C, Left antenna in ventral view. D, Left mesotarsomere 5 in ventral view. Scale bars: A = 1 mm, B = 0.5 mm, C = 0.25 mm, D = 0.125 mm.