Nemastoma bidentatum schmidti, Novak & Novak & Kozel & Schaider & Komposch & Lipovšek & Podlesnik & Paušič & Raspotnig, 2021

Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, European Journal of Taxonomy 777, pp. 1-67 : 47-49

publication ID

https://doi.org/ 10.5852/ejt.2021.777.1561

publication LSID

lsid:zoobank.org:pub:C86302D6-49F7-41A6-9BAC-94FDFED0D27B

DOI

https://doi.org/10.5281/zenodo.5637250

persistent identifier

https://treatment.plazi.org/id/389E4C27-24B9-44F9-A724-11475CB9E12B

taxon LSID

lsid:zoobank.org:act:389E4C27-24B9-44F9-A724-11475CB9E12B

treatment provided by

Felipe

scientific name

Nemastoma bidentatum schmidti
status

subsp. nov.

Nemastoma bidentatum schmidti ssp. nov. × N. bidentatum sparsum Gruber & Martens, 1968

Figs 3 View Fig , 9P View Fig , 10O View Fig

Nemastoma triste View in CoL – Roewer 1917: 149: [partim: Celje / Cilli, Maribor / Marburg].

Nemastoma bidentatum bidentatum View in CoL × N. bidentatum sparsum – Gruber & Martens 1968: 142, 147, figs 12–13 [partim: Novo mesto; Županja]. — Martens 1978: 107, fig 146 [partim: Županja; Novo mesto, Mt. Snežnik]. — Novak & Gruber 2000: 286 [partim: Novo mesto]. — Novak et al. 2002: 136 [partim: Ptuj]. — Novak 2004b: 246 [partim: Županja].

Nemastoma bidentatum View in CoL ssp. – Novak & Gruber 2000: 286 [partim: Celje, Maribor].

Diagnosis

Hybrids exhibiting varying Ch-Apo morphology, but with knee-like flexion of anterior margin, and either being relatively tiny with more or less elongated superior portion or more massive with superior portion drawn out in tiny apical portion, and club-shaped Pa-Fe with intermediate form with respect to both subspecies.

Material examined

SLOVENIA – VM91 • 2 ♂♂, 4 ♀♀; Čemšemik ; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1183/2002, rev. 2018); PMSL 1 ♂, 1 ♀; Jesenovo – Znojile; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1187/2002, TN rev. 2018); PMSL 6 ♂♂, 5 ♀♀; Mt. Čemšeniška planina; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1191/2002, rev. 2018); PMSL 2 ♀♀; Zaplanina, Ločica pri Vranskem ; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1210/2002, rev. 2018); PMSL 1 ♂, 5 ♀♀; ibid.; (1179/2002, rev. 2018); PMSL. – VM92 4 ♂♂, 9 ♀♀; Ločica pri Vranskem; 10 Aug. 1985; L. Slana Novak , M. Slana Novak and T. Novak leg. ( LSN 101 About LSN /1986, TN rev. 2018); PMSL. – WM01 7 ♂♂, 5 ♀♀; Marija Reka ; 21 Jul. 2002; L. Slana Novak and T. Novak leg. (1186/2002, rev. 2018); PMSL. – WM05 4 ♂♂, 7 ♀♀; Slovenj Gradec; 9 Oct. 1983; T. Novak and L. Slana Novak leg. ( LSN 113 About LSN /1983, TN rev. 2018); 3 ♂♂, 4 ♀♀; ibid.; 6 Jul. 2003; L. Slana Novak and T. Novak leg. (190/2003, rev. 2018); PMSL. – WM06 1 ♂, 2 ♀♀; Dravograd; 25 Oct. 1995; S. Hudrap and M. Pavlin leg. (11/1996, rev. 2018); PMSL. – WM10 4 ♂♂, 1 ♀; Lože – Globoko ; 14 Aug. 1983; T. Novak and L. Slana Novak leg. ( LSN 45 About LSN /1983, TN rev. 2018); PMSL. – WM11 1 ♂, 2 ♀♀; Brdce ; 22 Apr. 1995; S. Brelih leg. (758/1998, rev. 2018); PMSL 1 ♂, 3 ♀♀; Udmat ; 15 Aug. 1983; T. Novak and L. Slana Novak leg. ( LSN 47 About LSN /1983, TN rev. 2018); PMSL. – WM12 7 ♂♂, 3 ♀♀; Arja vas; 10 Jul. 1986; L. Slana Novak and T. Novak leg. (513/2002, rev. 2018); PMSL 1 ♂, 1 ♀; Velika Pirešica ; 10 Jul. 1986; L. Slana Novak and T. Novak leg. (452/2002, rev. 2018); PMSL 1 ♂, 4 juvs; ibid.; (470/2002, rev. 2018); PMSL. – WM13 4 ♂♂, 3 ♀♀; Vodosteč, Črnova ; 10 Jul. 1986; L. Slana Novak and T. Novak leg. (453/2002, rev. 2018); PMSL 2 ♂♂; ibid.; 22 Apr. 1995; S. Brelih leg. (772/1998, rev. 2018); PMSL 1 ♂, 1 ♀; ibid.; 21 Apr. 2002; L. Slana Novak and T. Novak leg. (737/2002, rev. 2019); PMSL. – WM16 2 ♂♂, 3 ♀♀; Sv. Trije Kralji, Muta; 20 Aug. 2005; L. Slana Novak and T. Novak leg. (79/2005, rev. 20018); PMSL. – WM20 2 ♂♂, 3 ♀♀; Jurklošter ; 15 Aug. 1983; L. Slana Novak and T. Novak leg. ( LSN 53 About LSN /1983, TN rev. 2018); PMSL. – WM21 1 ♂, 2 ♀♀; Lahovna ; 22 Apr. 1995; S. Brelih leg. (740/1998, TN rev. 2018); PMSL 2 ♂♂, 3 ♀♀, 1 juv.; Trobni dol; 16 Aug. 1983; T. Novak and L. Slana Novak leg. ( LSN 48 About LSN /1983, TN rev. 2018); PMSL. – WM22 Celje • 1 ♂; ibid.; 30 Oct. 2000; N. Žmaher leg. (881/2000, rev. 2018); PMSL 7 ♂♂, 2 ♀♀; Šmartinsko jezero; 25 Mar. 1995; S. Brelih leg. (704/1998, rev. 2018); PMSL 1 ♂, 4 ♀♀; Vojnik ; 17 Aug. 1983; T. Novak and L. Slana Novak leg. ( LSN 58 About LSN /1983, TN rev. 2018); PMSL. – WM23 1 ♂, 1 ♀; Črešnjice ; 25 Mar. 1995; S. Brelih leg. (709/1998, rev. 2018); PMSL 3 ♂♂, 3 ♀♀, l juv.; Frankolovo ; 17 Aug. 1983; L. Slana Novak and T. Novak leg. ( LSN 76 About LSN /1983, TN rev. 2018); PMSL 1 ♂; Mt. Konjiška gora; 14 Jun. 1981; V. Kuštor, T. Novak leg. (1120/1981, rev. 2018); PMSL 1 ♂, 2 ♀♀; Mt. Stenica ; 30 May 1981; T. Novak, V. Kuštor leg. (228/1981, rev. 2018); PMSL 1 ♀; Stenica ; 24 Jul. 2002; B. Drovenik and A. Gergeli leg. (1138/2018); PMSL 1 ♀; ibid.; 24 Jul.–21 Aug. 2002; B. Drovenik and A. Gergeli leg. ( PK 73 /2012, TN rev. 2018); PMSL 1 ♀; ibid.; ( PK 12 /2013, TN rev. 2018); PMSL 1 ♂; ibid.; 21 Aug. 2002; B. Drovenik and A. Gergeli leg. (1071/2018); PMSL 1 ♀; ibid.; 21 Aug.−16 Sep. 2002; B. Drovenik and A. Gergeli leg. (782/2018); PMSL 1 ♂, 1 ♀; ibid.; (819/2018); PMSL 1 ♀; ibid.; (856/2018); PMSL 1 ♂, 1 ♀; ibid.; (804/2018); PMSL 1 ♀; ibid.; 16 Sep.−14 Oct. 2002; B. Drovenik and A. Gergeli leg. (817/2018); PMSL 1 ♀; ibid.; 14 Oct. 2002; B. Drovenik and A. Gergeli leg. (361/ 2018); PMSL 1 ♀; ibid.; (783/2018); PMSL 1 ♂, 1 ♀; ibid.; (794/2018); PMSL 1 ♂, 1 ♀; ibid.; (814/2018); PMSL 1 ♀; ibid.; (824/2018); PMSL 1 ♀; ibid.; (831/2018); PMSL 1 ♀; ibid.; (846/2018); PMSL 1 ♂; ibid.; 11 Nov. 2002; B. Drovenik and A. Gergeli leg. (208/2018); PMSL 1 juv.; ibid.; (266/2018); PMSL 1 ♂; ibid.; 11 Nov. 2002 − 4 Mar. 2003; B. Drovenik and A. Gergeli leg. (694/2018); PMSL 1 ♀; ibid.; (698/2018); PMSL 2 ♀♀; ibid.; (701/2018); PMSL 1 ♂, 1 ♀; ibid.; (710/2018); PMSL 1 ♀; ibid.; (716/2018); PMSL 1 ♀; ibid.; (721/2018); PMSL 1 ♂; ibid.; (724/2018); PMSL 3 ♂♂, 2 ♀♀; ibid.; (729/2018); PMSL 1 ♂; ibid.; 8 Apr. 2003; B. Drovenik and A. Gergeli leg. (267/ 2018); PMSL 3 juvs; ibid.; (271/2018); PMSL 1 ♂; ibid.; 8 Apr. 2003; B. Drovenik and A. Gergeli leg. (189/ 2018); PMSL 1 juv.; ibid.; (266/2018); PMSL. – WM24 1 ♂, 1 ♀; Jurgovo ; 24 Jul.–3 Aug. 2005; A. Kapla leg. (178/2005, rev. 2018); PMSL 3 ♂♂; Resnik; 12 Aug. 1984; L. Slana Novak , M. Slana Novak and T. Novak leg. ( LSN 252 About LSN /1984, TN rev. 2018); PMSL. – WM25 3 ♂♂; Glažuta, Mislinjski jarek; 9 Sep. 2017; M. Delakorda , S. Lipovšek, L. Slana Novak and T. Novak leg. (107/2018); PMSL 1 ♀; ibid.; 19 Sep. 2018; S. Lipovšek , P. Kozel and T. Novak leg. (132/2018); PMSL 1 ♂; Ribnica na Pohorju; 4 Jul. 1939; J. Hadži leg. (145/1984, rev. 2018); PMSL. – WM30 1 ♂, 1 ♀; Loke – Lažiše ; 11 Jul. 2003; T. Novak leg. (154/2003, rev. 2018); PMSL. – WM31 3 ♂♂, 2 ♀♀; Bezovje pri Šentjurju ; 16 Aug. 1983; L. Slana Novak and T. Novak leg. ( LSN 56 About LSN /1983, TN rev. 2018); PMSL 1 ♂; Slivniško jezero; 7 Apr. 1995; S. Brelih leg. (701/1998, rev. 2018); PMSL. – WM33 3 ♂♂, 2 ♀♀, 1 juv.; Tepanje ; 17 Aug. 1983; T. Novak and L. Slana Novak leg. ( LSN 62 About LSN /1983, TN rev. 2018); PMSL 2 ♂♂; Žiče monastery; 25 Mar. 1995; S. Brelih leg. (706/1998, rev. 2018); PMSL. – WM40 1 ♂; Lesično ; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1130/2002, rev. 2018); PMSL. – WM41 1 ♂, 5 ♀♀; Podčetrtek; 13 Aug. 1983; L. Slana Novak and T. Novak leg. ( LSN 51 About LSN /1983, TN rev. 2018); PMSL 2 ♂♂, 3 ♀♀; ibid.; 7 Jul. 2012; T. Novak leg. (51/2012, rev. 2018); PMSL 2 ♂♂, 3 ♀♀; Virštajn ; 7 Apr. 1995; S. Brelih leg. (699/1998, rev. 2018); PMSL. – WM42 1 ♀; Mt. Boč ; 17 Apr. 1982; T. Novak and G. Kolar leg. (125/1982, rev. 2018); PMSL 2 ♂♂, 1 ♀; ibid.; 16 Oct. 1983; L. Slana Novak and T. Novak leg. ( LSN 118 About LSN /1983, TN rev. 2018); PMSL 2 ♂♂; Mestinje ; 13 Jul. 2002; L. Slana Novak and T. Novak leg. (1165/2002, rev. 2018); PMSL 1 ♂, 1 ♀; ibid.; (1168/2002, rev. 2018); PMSL 1 ♂, 1 ♀; Zgornji Gabrnik ; 11 Oct. 1997; L. Slana Novak and T. Novak leg. (412/1998, rev. 2018); PMSL. – WM43 4 ♂♂, 1 ♀; Ložnica ; 17 Aug. 1983; T. Novak and L. Slana Novak leg. ( LSN 75 About LSN /1983, TN rev. 2018); PMSL. – WM44 2 ♂♂, 4 ♀♀; Kočno pri Polskavi ; 8 Aug. 2015; L. Slana Novak and T. Novak leg. (110/2018); PMSL. – WM45 1 ♂; Limbuš ; 11 Oct. 2001; D. Kos leg. (226/2004, rev. 2018); PMSL. – WM52 1 ♀; Sv. Donat, Donačka gora; 16 Sep. 2002; B. Drovenik and A. Pirnat leg. (247/2018); PMSL 1 ♀; ibid.; (248/2018); PMSL 1 ♀; ibid.; 14 Oct. 2002; B. Drovenik and A. Pirnat leg. (1032/2018); PMSL 6 ♂♂, 2 ♀♀, 2 juvs; Tuncovec; 4 Aug. 1983; T. Novak , M. Štangelj and L. Slana Novak leg. ( LSN 66 About LSN /1983, TN rev. 20018); PMSL. – WM53 4 ♂♂, 1 ♀; Majšperk; 24 Jun. 2007; T. Novak leg. (43/2018); PMSL 4 ♂♂, 1 ♀; Pečke ; 10 Apr. 1995; S. Brelih leg. (713/1998, rev. 2018); PMSL 1 ♂; Stogovci ; 24 Jun. 2007; T. Novak leg. (50/2007, rev. 2018); PMSL. – WM62 1 ♂, 2 ♀♀; Dobovec pri Rogatcu ; 7 Oct. 2001; T. Novak leg. (605/2002, rev. 2007); PMSL 2 ♂♂; ibid.; 16 Aug. 2003; J. Ravljen and T. Novak leg. (807/2003, rev. 2018); PMSL 4 ♂♂; ibid.; 16 Aug. 2003; J. Ravljen , L. Slana Novak and T. Novak leg. (812/2003, rev. 2007); PMSL 1 ♂, 2 ♀♀; ibid.; 30 Sep. 2012; T. Novak leg. (63/2015); PMSL 7 ♂♂, 4 ♀♀, 1 juv.; Kozminci; 6 Aug. 1983; T. Novak , M. Štangelj and L. Slana Novak leg. ( LSN 61 About LSN /1983, TN rev. 2018); PMSL 1 ♀; Mt. Macelj ; Jun. 1980; V. Kuštor and T. Novak leg. ( LSN 5 About LSN /1983, TN rev. 2018); PMSL 2 ♀♀; Žale, Dobrina ; 24 Apr. 1998; S. Brelih leg. (85/2001, rev. 2018); PMSL. – WM63 4 ♂♂, 1 ♀; Majski vrh − Tržec; 6 Aug. 1983; T. Novak , M. Štangelj and L. Slana Novak leg. ( LSN 65 About LSN /1983, TN rev. 2018); PMSL. – WM64 3 ♂♂, 3 ♀♀; Ptuj; 6 Aug. 1983, T. Novak , M. Štangelj and L. Slana Novak leg. ( LSN 77 About LSN /1983, TN rev. 2018); PMSL. – WM65 2 ♂♂; Selca; 14 Jun. 1984; L. Slana Novak , M. Slana Novak and T. Novak leg. ( LSN 238 About LSN /1984, TN rev. 2018); PMSL 1 ♂, 2 ♀♀; Zgornja Voličina ; 23 Apr. 1998; S. Brelih leg. (124/2001, rev. 2018); PMSL. – WM73 1 ♂, 4 ♀♀; Berinjak ; 24 Jun. 2007; T. Novak leg. (36/2007, rev. 2018); PMSL 1 ♂, 8 ♀♀; Borl castle, Dolane; 7 Aug. 1983; L. Slana Novak , M. Štangelj and T. Novak leg. ( LSN 59 About LSN /1983, TN rev. 2018); PMSL 1 ♂, 2 ♀♀; Ljubslava , Podlehnik; 24 Jun. 2007; T. Novak leg. (49/2007, TN rev. 2018); PMSL 5 ♂♂; Spodje Gruškovje , Podlehnik; 24 Jun. 2007; T. Novak leg. (13/2007, rev. 2018); PMSL 4 ♂♂, 1 ♀; Vareja; 6 Aug. 1983; T. Novak , M. Štangelj and L. Slana Novak leg. ( LSN 78 About LSN /1983, TN rev. 2018); PMSL. – WM76 4 ♂♂, 3 ♀♀; Brengova ; 26 Jul. 1983; T. Novak and L. Slana Novak leg. ( LSN 63 About LSN /1983, TN rev. 2018); PMSL 2 ♂♂, 1 ♀; Kočki vrh; 30 Jun. 2002; T. Novak leg. (783/2002, rev. 2018); PMSL 1 ♂, 2 ♀♀; ibid.; (789/2002, rev. 2018); PMSL. – WM83 3 ♂♂, 5 ♀♀; Korenjak – Turški vrh; 7 Aug. 1983; T. Novak , M. Štangelj and L. Slana Novak leg. ( LSN 55 About LSN /1983, TN rev. 2018); PMSL. – WM84 4 ♂♂, 3 ♀♀; Vudina, Velika Nedelja ; 29 May 2001; P. Igerc and T. Novak leg. (22/2001, rev. 2018); PMSL. – WM85 2 ♀♀; Radoslavci; 10 Sep. 1983; T. Novak , M. Slana Novak and L. Slana Novak leg. ( LSN 117 About LSN /1983, TN rev. 2018); PMSL. – WM94 2 ♂♂, 3 ♀♀; Pavlovci ; 2 Apr. 1983; T. Novak and L. Slana Novak leg. ( LSN 6 About LSN /1983, TN rev. 2018); PMSL. – WM95 Noršinci pri Ljutomeru ; 27 Aug. 1983; T. Novak and L. Slana Novak leg. 2 ♂♂, 2 ♀♀ ( LSN 109 About LSN /1983, TN rev. 2018); PMSL. – WM95 3 ♂♂, 3 ♀♀; Gresovščak; L. Slana Novak and T. Novak leg. (163/2012, rev. 2018); PMSL 2 ♂♂, 2 ♀♀; Slamnjak , Ljutomer; 10 Jun. 1984; L. Slana Novak and T. Novak leg. ( LSN 241 About LSN /1984, rev. 2018); PMSL .

Remarks

To date, N. b. schmidti ssp. nov. has been reported under N. b. bidentatum , and N. b. gruberi ssp. nov. under N. b. sparsum. Since hybrids N. b. bidentatum × N. b. sparsum have not been identified in any locality, all these published data likely refer either to N. b. schmidti ssp. nov. × N. b. sparsum or N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. Nemastoma b. bidentatum referred from Germany ( Schönhofer & Holle 2007) is a hybrid, probably N. b. schmidti ssp. nov. × N. b. sparsum. Since both N. b. sparsum and N. b. gruberi ssp. nov. inhabit the lowland in Croatia, hybrids south of the Sava River or nearby probably belong to N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov.

PMSL

Slovenia, Ljubljana, Slovenian Museum of Natural History

L

Nationaal Herbarium Nederland, Leiden University branch

T

Tavera, Department of Geology and Geophysics

PMSL

Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)

M

Botanische Staatssammlung München

S

Department of Botany, Swedish Museum of Natural History

N

Nanjing University

V

Royal British Columbia Museum - Herbarium

B

Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet

A

Harvard University - Arnold Arboretum

P

Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants

J

University of the Witwatersrand

G

Conservatoire et Jardin botaniques de la Ville de Genève

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Opiliones

Family

Nemastomatidae

Genus

Nemastoma

Loc

Nemastoma bidentatum schmidti

Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther 2021
2021
Loc

Nemastoma bidentatum

Novak T. & Gruber J. 2000: 286
2000
Loc

Nemastoma bidentatum bidentatum

Novak T. 2004: 246
Novak T. & Slana L. & Cervek N. & Mlakar M. & Zmaher N. & Gruber J. 2002: 136
Novak T. & Gruber J. 2000: 286
Martens J. 1978: 107
Gruber J. & Martens J. 1968: 142
1968
Loc

Nemastoma triste

Roewer C. F. 1917: 149
1917
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