Phylloicus iridescens Banks, 1941

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 61-62

Phylloicus iridescens Banks

Figs. 59 , 60 , 111

Phylloicus iridescens Banks, 1941:397 [Type locality: Dominican Republic, Constanza to V. Nuevo; MCZ; male]. — Flint 1967:18 [lectotype male].

This species is most similar to P. cubanus , pulchrus , and superbus . In P. iridescens the orange bands of the forewing are narrower ( Fig. 111 ) than in the other three species, and the male genitalia are distinguished by a tapering harpago.

Adult. Forewing length 8.1­9.5 mm, n = 12.

Head dark brown. Maxillary palps dark brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax dark brown; ventrolateral thorax chestnut brown. Femora golden; foretibiae dark brown; mesotibiae dark brown; metatibiae dark brown; foretarsi dark brown; mesotarsi white; metatarsi dark brown. Metathoracic leg of male without posterior fringe. Tibial spur formula 2,4,2. Forewing flat; dark brown; with two transverse bands; proximal band orange, reaching posterior wing margin, at least 1/2 width of wing; distal band orange, beginning at anterior wing margin, at least 1/2 width of wing; basal cells without setae, membrane iridescent ( Fig. 111 ). Hind wing basal brush absent.

Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VII with short, acute anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX with mesal ridge extending full length of segment; posterior margin continuous with basodorsal process of tergum X ( Fig. 59B ); lateral ridge present; dorsal pleural setae approximately 15, ventral pleural setae absent ( Fig. 59A ); sternum IX with paired mesolateral ridges; sternum IX ( Fig. 59C ). Preanal appendage less than 2/3 length of tergum X, narrowly elliptic, setae long, but not filamentous or longer than appendage ( Fig. 59A, B ). Tergum X without basal lobes; basodorsal process short and digitate; basolateral processes absent; apex, in lateral view, rounded, in dorsal view, notched, notch shallow ( Fig. 59A, B ). Harpago rounded; peglike setae few, apical ( Fig. 59A, C ). Phallic endotheca with paired apicolateral lobes, apicolateral lobes rounded and with digitate apical lobe; phallotremal sclerites very large, longest dimension twice diameter of phallobase; dorsal sclerite two­armed, in lateral view U­shaped ( Fig. 59D, E ).

Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII cleft posteromesally to anterior ridge; sternum VIII ( Fig. 60C ). Tergum IX with mesal ridge extending 1/3 length of segment ( Fig. 60B ). Sternum IX anterior lobes smooth and indistinct, posterior lobes striate, without distinct area of thin cuticle or invagination ( Fig. 60A ). Tergum X appendage longer than mesal lobe, base distinct, apex rounded; mesal lobe lightly sclerotized; digitate lateral processes absent ( Fig. 60B ). Sternum X without setae in membrane ( Fig. 60A ). Vaginal apparatus posterior sclerite elongate; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite triangular; posterior end of spermatheca a sclerotized cone ( Fig. 60A ).

Material examined. DOMINICAN REPUBLIC: 20 km S Constanza, 3­7.vi.1969, Flint & Gomez — 3 males, 2 females ( NMNH) ; Constanza to V. Nuevo , 3000 ­ 7000 f m, 1.viii.1938, Darlington — lectotype male, 1 male, 1 female paralectotypes ( MCZ) ; Loma Rucilla & mts., 5000­8000 f m, 1.vi.1938, Darlington — 1 male paralectotype ( MCZ) ; Valle Nuevo SE Constanza , 2134 m, 1.viii.1938, Darlington — 2 females ( MCZ) ; Dajabon: 13 km S Loma de Cabrera , 400 m, 20­22.v.1973, D & M Davis — 3 males, 2 females ( NMNH) .

Distribution. Dominican Republic.

Figures

FIGURE 59. Phylloicus iridescens. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view.

FIGURE 60. Phylloicus iridescens. Female (UMSP000009755): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view.

FIGURES 109-112. — 109. Phylloicus elegans (composite of UMSP000000172 and UMSP000022365). Forewing. — 110. Phylloicus flinti (UMSP000010104). Forewing. — 111. Phylloicus iridescens (composite of UMSP000009750 and UMSP000009751). Forewing. — 112. Phylloicus paprockii (composite of UMSP000068390-2). Forewing.