Phylloicus quadridigitatus, Prather , 2003

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 91-92

Phylloicus quadridigitatus , new species

Fig. 96

This species is known only from a single male, with antennae and wings badly rubbed. Thus, any ornamentation of wings or antennae is unknown. The phallic endotheca is not fully everted, so I am unable to describe the membranous lobes. Phylloicus quadridigitatus is distinguished by the two pairs of digitate processes of tergum X, from which it takes its name.

Adult. Forewing length 7.3 mm, n = 1.

Head black, including setal warts. Maxillary palps black. Antenna twice forewing length; dark brown. Prothorax black; dorsal pterothorax black; ventrolateral thorax chestnut brown. Femora chestnut brown; tibiae golden brown; tarsi golden brown. Metathoracic leg of male with posterior fringe of long setae. Tibial spur formula 2,2,2. Forewing flat; dark brown; without colored markings. Hind wing basal brush present in male.

Male. Preterminalic abdominal terga without anteromesal notches. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VII without anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX with paired mesal ridges extending length of segment; posterior margin obtuse ( Fig. 96B ); lateral ridge absent; dorsal pleural setae approximately 10, ventral pleural setae absent ( Fig. 96A ); sternum IX without mesolateral ridges; sternum IX ( Fig. 96C ). Preanal appendage less than 2/3 length of tergum X, widest near base, setae long, but not filamentous or longer than appendage ( Fig. 96A, B ). Tergum X without basal lobes; basodorsal process absent; basolateral processes long, length at least twice diameter; apex, in lateral view, with long digitate posterodorsal projection and short ventral projection, in dorsal view, cleft; with paired spine­like mesal setae at mid­length; posteriorly with two pairs of digitate processes, dorsomesal processes twice length of ventrolateral processes ( Fig. 96A, B ). Harpago large, rounded; peglike setae many, apical ( Fig. 96A, C ). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 96D, E ).

Female. Unknown.

Holotype male: BRAZIL: São Paulo: Alto da Serra , 29­30.x.1927, Zerny ( NMW).

Distribution. Brazil.

Figures

FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view.