Phylloicus panamensis, Prather, 2003

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 80-81

Phylloicus panamensis , new species

Figs. 80 , 81

Phylloicus panamensis is very similar to P. aeneus , nigripennis , and mexicanus . Like P. aeneus and mexicanus¸ the females of P. panamensis have a brush of setae laterally on tergum VIII. However, these specimens lack the coremata of P. aeneus , the orange diamond on the head distinguishing P. mexicanus , or the papillate terga II and III of P. nigripennis males. All the specimens I examined were preserved in ethanol, so details of wing color pattern or body coloring are unavailable.

Adult. Forewing length 12­13.5 mm, n = 41.

Head golden brown. Maxillary palps golden brown. Antenna twice forewing length; golden brown. Prothorax golden brown; dorsal pterothorax golden brown; ventrolateral thorax golden brown. Legs golden brown. Tibial spur formula 2,4,3. Forewing apical third folded obliquely toward midline; golden brown; without colored markings. Hind wing basal brush present in female.

Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VII with short, acute anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX with short mesal ridge extending from anterior notch; posterior margin irregular ( Fig. 80B ); lateral ridge present; dorsal pleural setae approximately 8, ventral pleural setae approximately 5 ( Fig. 80A ); sternum IX with paired mesolateral ridges; sternum IX ( Fig. 80C ). Preanal appendage approximately length of tergum X, of uniform diameter throughout length, setae long, but not filamentous or longer than appendage ( Fig. 80A, B ). Tergum X sagittate basally; basodorsal process absent; basolateral processes absent; apex, in lateral view, rounded, in dorsal view, notched, notch triangular; dorsal surface setose; with anterodorsal lateral processes ( Fig. 80A, B ). Harpago long, curving mesally and tapering apically; peglike setae few, apical ( Fig. 80A, C ). Phallic endotheca with paired basolateral lobes, basolateral lobes large and round and with digitate apical lobe; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite two­armed, in lateral view U­shaped ( Fig. 80D, E ).

Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII with dense posterolateral brush of setae; Fig. 81B ; sternum VIII cleft posteromesally to anterior ridge; sternum VIII ( Fig. 81C ). Tergum IX with mesal ridge extending 1/3 length of segment ( Fig. 81B ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, punctate, with shallow pockets lateral to vaginal opening ( Fig. 81A ). Tergum X appendage longer than mesal lobe, base marked by faint suture line, apex triangular; mesal lobe lightly sclerotized; digitate lateral processes absent ( Fig. 81B ). Sternum X without setae in membrane ( Fig. 81A ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite truncate anteriorly, posterolateral projections acute; posterior sclerite ovoid; posterior end of spermatheca a sclerotized sphere ( Fig. 81A ).

Holotype male: PANAMA: Chiriqui: Guadalupe Arriba , 08°52'26”N, 82°33'13"W,. iv.1984, Wolda ( NMNH).

Paratypes: COSTA RICA: Puntarenas: Río Bellavista , ca. 1.5 km NW Las Alturas, 08°57'04”N, 82°50'46"W, 1400 m, 8­9.iv.1987, Holzenthal, Hamilton, & Heyn — 1 male ( UMSP) ; PANAMA: Chiriqui: Guadalupe Arriba , 08°52'26”N, 82°33'13"W,. iv.1984, Wolda — 4 females, 2 males ( NMNH) ; v.1984, Wolda — 3 females, 9 males ( UMSP) ; 8­ 14.viii.1984, Wolda — 1 female, 1 male ( UMSP) ;. v.1985, Wolda — 1 female, 2 males ( UMSP) ; 14 Aug­ 10.ix.1985, Wolda — 3 males ( NMNH) ; 25 Sep­ 15.x.1985, Wolda — 2 females, 2 males ( NMNH) .

Distribution. Costa Rica, Panama.

Etymology. Named for the country in which the types were collected.

Figures

FIGURE 80. Phylloicus panamensis. Male (UMSP000018646): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view.

FIGURE 81. Phylloicus panamensis. Female (UMSP000041338): A—sterna IX, X and vaginal apparatus, ventral view; B—terga VIII­X, dorsal view; C—sternum VIII, ventral view.