Adeliini Viereck, 1918
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publication ID |
https://doi.org/ 10.11646/zootaxa.4926.1.1 |
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publication LSID |
lsid:zoobank.org:pub:F151B4E4-26FD-42CE-9F4D-0D8931AAF93F |
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DOI |
https://doi.org/10.5281/zenodo.4500472 |
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persistent identifier |
https://treatment.plazi.org/id/5466E313-FFA4-511B-519C-FB1EFE0DFBD6 |
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treatment provided by |
Plazi |
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scientific name |
Adeliini Viereck, 1918 |
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Tribe Adeliini Viereck, 1918
Diagnosis. Head transverse, sculptured ( Figs 1B View FIGURE 1 , 3D View FIGURE 3 , 5B View FIGURE 5 , 7B View FIGURE 7 , 9C View FIGURE 9 , 11K View FIGURE 11 , 12I View FIGURE 12 , 13B View FIGURE 13 , 15C View FIGURE 15 , 17B View FIGURE 17 ); ocelli small; frons flat or concave medially, with or without midlongitudinal groove ( Figs 1C View FIGURE 1 , 4C View FIGURE 4 , 5C View FIGURE 5 , 7C View FIGURE 7 , 9D View FIGURE 9 , 11C View FIGURE 11 , 12C View FIGURE 12 , 13C View FIGURE 13 , 15D View FIGURE 15 , 17C View FIGURE 17 ); eyes setose ( Figs 1B View FIGURE 1 , 3D View FIGURE 3 , 5B View FIGURE 5 , 7B View FIGURE 7 , 12I View FIGURE 12 , 13B View FIGURE 13 , 15C View FIGURE 15 , 17B View FIGURE 17 ) or glabrous ( Figs 9C View FIGURE 9 , 11K View FIGURE 11 ); malar suture complete and deep ( Figs 1E View FIGURE 1 , 3D View FIGURE 3 , 5D & E View FIGURE 5 , 7D View FIGURE 7 , 10B & E View FIGURE 10 , 11I & K View FIGURE 11 , 15E View FIGURE 15 ); occipital carina often complete, joining hypostomal carina ventrally ( Figs 4A View FIGURE 4 , 6A View FIGURE 6 , 8A View FIGURE 8 , 10A View FIGURE 10 , 14A View FIGURE 14 , 16A View FIGURE 16 ), rarely absent ventrally ( Figs 1A & E View FIGURE 1 ) (in Carinadelius gen. nov.) without joining hypostomal carina; palps short and thick, maxillary palp often 5-segmented ( Figs 1E View FIGURE 1 , 4A View FIGURE 4 , 13E View FIGURE 13 ) rarely 3-segmented ( Fig. 12F View FIGURE 12 ) (in Paradelius ), labial palp 3-segmented; antenna more or less thickened, 20–22 antennomeres; scape rather long, wide; pedicel short; first flagellomere longer than second; subapical flagellomeres of female quadrate in several species ( Figs 2A View FIGURE 2 , 5A View FIGURE 5 , 7C View FIGURE 7 ); tentorial pits deep ( Figs 1B View FIGURE 1 , 3D View FIGURE 3 , 5B View FIGURE 5 , 7B View FIGURE 7 , 9C View FIGURE 9 , 11K View FIGURE 11 , 12I View FIGURE 12 , 13B View FIGURE 13 , 15C View FIGURE 15 , 17B View FIGURE 17 ); clypeus smooth or sculptured ( Figs 1B View FIGURE 1 , 3D View FIGURE 3 , 5B View FIGURE 5 , 7B View FIGURE 7 , 9C View FIGURE 9 , 11K View FIGURE 11 , 12I View FIGURE 12 , 13B View FIGURE 13 , 15C View FIGURE 15 , 17B View FIGURE 17 ); labrum concealed by clypeus ( Figs 1B View FIGURE 1 , 3D View FIGURE 3 , 5B View FIGURE 5 , 7B View FIGURE 7 , 9C View FIGURE 9 , 11K View FIGURE 11 , 12I View FIGURE 12 , 13B View FIGURE 13 , 15C View FIGURE 15 , 17B View FIGURE 17 ); mesosoma often dorso-ventrally flattened ( Figs 1E View FIGURE 1 , 4B View FIGURE 4 , 6B View FIGURE 6 , 7F View FIGURE 7 , 10B View FIGURE 10 , 11I View FIGURE 11 , 13E View FIGURE 13 , 15E View FIGURE 15 , 17E View FIGURE 17 ); propleuron short, crenulate posteriorly ( Figs 1E View FIGURE 1 , 4B View FIGURE 4 , 6B View FIGURE 6 , 7F View FIGURE 7 , 10B View FIGURE 10 , 11I View FIGURE 11 , 13E View FIGURE 13 , 15E View FIGURE 15 , 17E View FIGURE 17 ); propleural lobe distinct or indistinct ( Figs 1E View FIGURE 1 , 4B View FIGURE 4 , 6B View FIGURE 6 , 7F View FIGURE 7 , 10B View FIGURE 10 , 11I View FIGURE 11 , 13E View FIGURE 13 , 15E View FIGURE 15 , 17E View FIGURE 17 ); mesoscutum smooth to sculptured ( Figs 1D View FIGURE 1 , 4C View FIGURE 4 , 5F View FIGURE 5 , 7E View FIGURE 7 , 9F View FIGURE 9 , 11L View FIGURE 11 , 12B View FIGURE 12 , 13D View FIGURE 13 , 15D View FIGURE 15 , 17C View FIGURE 17 ); notauli absent ( Figs 1D View FIGURE 1 , 4C View FIGURE 4 , 5F View FIGURE 5 , 7E View FIGURE 7 , 9F View FIGURE 9 , 11L View FIGURE 11 , 12B View FIGURE 12 , 13D View FIGURE 13 , 15D View FIGURE 15 , 17C View FIGURE 17 ); prepectal carina often absent ( Figs 1E View FIGURE 1 , 4B View FIGURE 4 , 6B View FIGURE 6 , 7F View FIGURE 7 , 10B View FIGURE 10 , 11I View FIGURE 11 , 13E View FIGURE 13 , 15E View FIGURE 15 ), rarely present ( Fig. 12H View FIGURE 12 , 17E View FIGURE 17 ) (in Sinadelius and Paradelius ); postpectal carina absent ( Fig. 9E View FIGURE 9 ); scutellar sulcus narrow, crenulated ( Figs 1D View FIGURE 1 , 4C View FIGURE 4 , 5F View FIGURE 5 , 7E View FIGURE 7 , 9F View FIGURE 9 , 11L View FIGURE 11 , 12B View FIGURE 12 , 13D View FIGURE 13 , 15D View FIGURE 15 , 17C View FIGURE 17 ); mesopleuron convex, smooth or sculptured ( Figs 1E View FIGURE 1 , 4B View FIGURE 4 , 6B View FIGURE 6 , 7F View FIGURE 7 , 10B View FIGURE 10 , 11I View FIGURE 11 , 12H View FIGURE 12 , 13E View FIGURE 13 , 15E View FIGURE 15 , 17E View FIGURE 17 ); precoxal sulcus smooth or crenulate entirely ( Figs 1E View FIGURE 1 , 4B View FIGURE 4 , 6B View FIGURE 6 , 7F View FIGURE 7 , 10B View FIGURE 10 , 11I View FIGURE 11 , 12H View FIGURE 12 , 13E View FIGURE 13 , 15E View FIGURE 15 , 17E View FIGURE 17 ); metapleuron smooth ( Figs 1E View FIGURE 1 , 7F View FIGURE 7 , 10B View FIGURE 10 , 13E View FIGURE 13 , 17E View FIGURE 17 ) or rugose ( Figs 4B View FIGURE 4 , 6B View FIGURE 6 , 15E View FIGURE 15 ); propodeum smooth or sculptured, often areolate, either divided into distinct anterior and posterior part or not ( Figs 1F View FIGURE 1 , 4D & E View FIGURE 4 , 6C View FIGURE 6 , 8B View FIGURE 8 , 10C & D View FIGURE 10 , 11D View FIGURE 11 , 12B View FIGURE 12 , 13F View FIGURE 13 , 14B View FIGURE 14 , 16B View FIGURE 16 , 17F View FIGURE 17 , 18A View FIGURE 18 ), rarely with distinct midlongitudinal carina and vertical lateral carina posteriorly ( Fig. 1F View FIGURE 1 ) (in Carinadelius gen. nov.); propodeal spiracles round; fore wing hyaline ( Figs 7A View FIGURE 7 , 10E View FIGURE 10 , 11A View FIGURE 11 , 12A View FIGURE 12 , 14C View FIGURE 14 , 18C View FIGURE 18 ) or infuscated ( Figs 2C View FIGURE 2 , 4F View FIGURE 4 , 6D View FIGURE 6 , 16C View FIGURE 16 ); pterostigma wide; vein 1-R1 distinct, long ( Figs 2C View FIGURE 2 , 4F View FIGURE 4 , 6D View FIGURE 6 , 10E View FIGURE 10 , 12A View FIGURE 12 , 14C View FIGURE 14 , 18C View FIGURE 18 ) to nearly absent ( Figs 7A View FIGURE 7 , 9A View FIGURE 9 , 11A View FIGURE 11 ); fore wing vein r present ( Figs 7A View FIGURE 7 , 9A View FIGURE 9 , 11B & F View FIGURE 11 ) or absent ( Figs 2C View FIGURE 2 , 4F View FIGURE 4 , 6D View FIGURE 6 , 12A View FIGURE 12 , 14C View FIGURE 14 , 18C View FIGURE 18 ); vein r-m absent ( Figs 2C View FIGURE 2 , 4F View FIGURE 4 , 6D View FIGURE 6 , 7A View FIGURE 7 , 10E View FIGURE 10 , 11A View FIGURE 11 , 12A View FIGURE 12 , 14C View FIGURE 14 , 16C View FIGURE 16 , 18C View FIGURE 18 ); vein SR1 not reaching wing margin; vein 1SR+M either connected with parastigma or 1-M; vein m-cu often postfurcal or interstitial ( Figs 2C View FIGURE 2 , 4F View FIGURE 4 , 6D View FIGURE 6 , 7A View FIGURE 7 , 10E View FIGURE 10 , 11A View FIGURE 11 , 12A View FIGURE 12 , 14C View FIGURE 14 , 16C View FIGURE 16 , 18C View FIGURE 18 ); vein M+CU straight or distinctly curved ( Figs 2C View FIGURE 2 , 4F View FIGURE 4 , 6D View FIGURE 6 , 7A View FIGURE 7 , 10E View FIGURE 10 , 11A View FIGURE 11 , 12A View FIGURE 12 , 14C View FIGURE 14 , 16C View FIGURE 16 , 18C View FIGURE 18 ); vein cu-a antefurcal, interstitial or postfurcal ( Figs 2C View FIGURE 2 , 4F View FIGURE 4 , 6D View FIGURE 6 , 7A View FIGURE 7 , 10E View FIGURE 10 , 11A View FIGURE 11 , 12A View FIGURE 12 , 14C View FIGURE 14 , 16C View FIGURE 16 , 18C View FIGURE 18 ); subdiscal cell open; hind wing with 3 hamuli; basal cell of hind wing narrow; mid coxa with transverse groove dorsally ( Figs 4B View FIGURE 4 , 6B View FIGURE 6 , 7F View FIGURE 7 , 16D View FIGURE 16 ); hind coxa smooth or rugose; hind femur and tibia flattened; hind tibial spurs long, inner spur approx. 0.5 × as long as basitarsus; hind basitarsus long, 0.8–0.9 × as long as combined length of 2 nd– 5 th tarsomeres; tarsal claws short, simple ( Fig. 12E View FIGURE 12 ); metasoma smooth or sculptured ( Figs 2B View FIGURE 2 , 4E View FIGURE 4 , 6C View FIGURE 6 , 8B View FIGURE 8 , 10D View FIGURE 10 , 11D View FIGURE 11 , 12G View FIGURE 12 , 14B View FIGURE 14 , 16B View FIGURE 16 , 18A View FIGURE 18 ), often compressed dorso-ventrally; metasomal tergites 1–3 formed into carapace, smooth or rugose; first and second metasomal suture present ( Figs 12G View FIGURE 12 , 14B View FIGURE 14 , 16B View FIGURE 16 , 18A View FIGURE 18 ) or absent ( Figs 2B View FIGURE 2 , 4E View FIGURE 4 , 6C View FIGURE 6 , 8B View FIGURE 8 , 10D View FIGURE 10 , 11D View FIGURE 11 , 16B View FIGURE 16 ); spiracle of metasomal tergites 6–7 absent; ovipositor short, always less than 0.25 × as long as hind tibia.
Genera included. Adelius Haliday , Carinadelius Ranjith & van Achterberg , gen. nov., Myriola Shestakov reinstated, Paradelius de Saeger , Sinadelius He & Chen , and Sculptomyriola Belokobylskij.
Distribution. Cosmopolitan.
Biology. Solitary endoparasitoids of leaf-mining Nepticulidae .
Comments. Shimbori et al. (2019) discussed possible apomorphic and synapomorphic character states between Cheloninae and Adeliini . Apart from the synapomorphic characters, the number of antennomeres is considered as plesiomorphic character to some extent (E.M. Shimbori, personal communication). Within the tribe it has been also found ( Kittel & Austin 2014; Kittel et al. 2015, 2016) that the division of the propodeum into distinct anterior and posterior parts (in Adelius , Sinadelius and Sculptomyriola ) is a plesiomorphic character as it is absent in other genera (in Carinadelius gen. nov., Myriola and Paradelius ). It is suggested that the new genus Carinadelius is probably derived from the genus Adelius .
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