Spelaeodiscus unidentatus unidentatus Bole, 1961

Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander & Feher, Zoltan, 2018, Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae), ZooKeys 769, pp. 13-48 : 13

Spelaeodiscus unidentatus unidentatus Bole, 1961 Figure 13

Spelaeodiscus unidentatus Bole, 1961: 205 fig. 1a-d.

Spelaeodiscus unidentatus - Bole 1965: 350-351, fig. 1a, 2b, plate 76, fig. C.

Spelaeodiscus (Spelaeodiscus) unidentatus - Gittenberger 1969: 297, fig. 1.

Spelaeodiscus unidentatus - Welter-Schultes 2012: 214.

Type material.

Obodska pećina, Rij. Crnojevića, Mtg., 1956 September, MZBI 1019/6 adult syntypes and some juvenile syntypes (photos of the sample where examined, Figure 12B ). See also Remarks.

Other material.

Bioče, N Titograd, Mtg. 23.09.1978, MZBI 15443 (photos of the sample where examined); Megara , Toluši, Titograd , Mtg. MZBI 2335 (photos of the sample where examined); Pećina od Zavora , Peuta , Titograd , Mtg. 01.11.1963, MZBI 2338 (photos of the sample where examined); Höhle bei Virpazar , leg. Dabović, coll. Edlauer, NHMW 48881/1; Brosine cave (33), leg. Kuscer 10266/67, coll. Edlauer, NHMW 38874/a (20 juvenile shells, "schlechte Expl "); Same data, NHMW 58874/4 ("bessere Expl "); Virpazar , mit "oculus mundi", leg. Dabović, coll. Edlauer, NHMW 78988/ 3 juvenile shells (det. Gittenberger, but identification not certain); Smarjetna gora + Grabocica cave (drawing of a cave on the label), ( Trnovo ), NHMW 78989/7; Same data, NHMW 48086/5; Vodna jama ( Lovćen), coll. Edlauer ex coll. Kuščer, NHMW 48544/1; Same locality, coll. Edlauer, NHMW 49764/1 (labelled as " Spelaeodiscus albanicus hadzii "); Montenegro, NNW of Virpazar, road between Rijeka Crnojevića and Virpazar, 0.4 km (in a straight line) W of Poseljani, 130 m a.s.l., 42°18.361'N, 19°02.898'E (site code: 20171020F), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 20.10.2017, DT/ca. 15, EZP/ca. 15, HA/ca. 15, HNHM 103248/5, PGB/ca. 15; Montenegro, W of Rijeka Crnojevića, 1 km (in a straight line) NNE of Zaćir, above Obodska pećina, near the road, 320 m a.s.l., 42°21.168'N, 19°00.125'E (site code: 2017.10.20 J), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 20.10.2017, DT/ca. 18, EZP/ca. 18, HA/ca. 18, HNHM 103249/5, PGB/ca. 18; Montenegro, 2 km north of Velje Brdo in the Zeta valley , north of Podgorica , 70 m a.s.l., 42°28.932'N, 19°14.454'E, leg. A. Reischütz, P. L. Reischütz & N. Steiner-Reischütz, May 2016, REI/5; Montenegro, Moraca valley , after the exit to the " China Road and Bridge" camp N of Bioće, 70 m a.s.l., 42°29.167'N, 19°18.646'E, leg. A. Reischütz, P. L. Reischütz & N. Steiner-Reischütz, May 2016, NHMW 112368/1 (photographed shell, Fig. 13G-L ), REI/10; Montenegro, road between Rijeka Crnojevića and Hrvasi, 180 m a.s.l., 42°21.491'N, 19°4.281'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Apr. 2012, REI/11; Montenegro, scree slope at the BMS-petrol station, south of the Morača gorge, 5 km south of Bioće, 90 m a.s.l., 42°29.064'N, 19°18.279'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Jul. 2008, REI/1; Montenegro, spring 1 km south of the junction towards Njive, south of Rijeka Crnojevića, 150 m a.s.l., 42°18.266'N, 19°2.900'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Mar. 2011, NHMW 112369/1 (photographed shell, Fig. 13M-R ); Montenegro, Žabljak Crnojevića, fortress, 50 m a.s.l., 42°19.030'N, 19°9.401'E (site code: 2015/007), leg. T. Deli, Z.P. Erőss & Z. Fehér, 25.05.2015, HNHM 103250/ 1 adult and several juvenile shells; Montenegro, Začir, “Pečina u pecko brdo" (cave), 450 m a.s.l., leg. L. Pintér, P. Subai, A. Szigeti, 20 Jul. 1972, NMBE 542072/76; Montenegro, Pečina u peckom, Brdu-bg Zacirnal ( Brdu-cave at Začir), leg. L. Pintér & P. Subai, 20 Jul. 1972, HNHM 65187/7 (one of them is photographed: Figs 13A-F ); Montenegro, Pečina u peckom Brdu bei Začir ( Brdu-cave at Začir), leg. H. Schütt, 06.06.1978, coll. Gy. Kovács, HNHM 65188/1; Same data, HNHM 42401/2; Montenegro, Rijeka Crnojevica , Obodska cave , 110 m a.s.l., 42°21.120'N, 19°0.318'E, leg. P. Subai & M. Szekeres, 23 Sep. 2006; NMBE 542071/1; Montenegro, Poseljani, on the Virpazar-Rijeka Crnojevića road, 150 m a.s.l., 42°18.347'N, 19°2.884'E (site code: 2015/016), leg. T. Deli, Z.P. Erőss & Z. Fehér, 25.05.2015, HNHM 103251/8; NHMW 112370/8, DT/8 .

Diagnosis.

Basal tooth/thickening low, not pointed; palatal part of peristome with strong incision at the position of the palatal tooth, palatal region strongly “pushed” from outside.

Description.

Spire elevated; protoconch consists of 1.5 whorls, roughly granulated/ ”hammered”, matte, not glossy; teleoconch with strong but dense, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (74-118 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75-4.5 whorls; aperture semilunar or triangular due to straight basal part; peristome thickened, slightly expanded and slightly reflected on the palatal, basal and umbilical areas; palatal tooth strong, pointed; palatal region of peristome with strong outer incision (i.e. the position of the palatal tooth is indicated with a groove on the outer side); basal portion of peristome straight, thickened; occasionally two small denticles visible, sometimes only the one situated closer to the palatal side is developed as a low denticle; umbilicus funnel-shaped, wide to relatively narrow (width depends on spire height).

Measurements. SW: 2.4-3.2 mm (median = 2.7 mm), SH: 1.3-1.8 mm (median = 1.5 mm), AW: 0.9-1.1 mm (median = 1.0 mm), AH: 1.0-1.4 mm (median = 1.1 mm), (n = 11; largest and smallest specimens of multiple populations measured).

Differential diagnosis.

See under S. unidentatus acutus ssp. n.

Variation among specimens.

This subspecies is particularly variable in terms of shell size. The shape of the thickening of the basal part of the peristome is also slightly variable, although it never develops to a pointed tooth.

Distribution.

This taxon is found north and northwest of the Shkodër Lake Basin in Montenegro (Figure 7 ). Albanian records given by Reischütz et al. (2013) actually refer to S. unidentatus acutus ssp. n.

Remarks.

The only sample in the MZBI, which was collected before the original description is from Obodska pećina represent the type sample of this species (MZBI 1019). Bole collected both S. obodensis and S. unidentatus at Obodska pećina in September, 1956, but described only one ( S. unidentatus ) in 1961, and the other ( S. obodensis ) in 1965.

Formerly, Peuta Cave population was incorrectly referred to as S. albanicus and was taken into consideration in that species’ Red List assessment ( Reischütz and Fehér 2017).

Figures

Figure 7. Distribution of Spelaeodiscus Brusina, 1886 species.

Figure 12. Type sample of Spelaeodiscus obodensis Bole, 1965 (A) and Spelaeodiscus unidentatus unidentatus Bole, 1961 (B).

Figure 13. Spelaeodiscus unidentatus unidentatus Bole, 1961. A-F Pecina u Peckom Brdu cave, above Zacir (HNHM 65187, topotypic specimen) G-L Montenegro, Moraca valley, after the exit to the " China Road and Bridge " camp N of Bioce (NHMW 112368) M-R Montenegro, spring 1 km south of the junction towards Njive (NHMW 112369).