Exocelina polita (Sharp, 1882)

Shaverdo, Helena V., Surbakti, Suriani, Hendrich, Lars & Balke, Michael, 2012, Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 250, pp. 1-76 : 26-27

19. Exocelina polita (Sharp, 1882) Figs 11 A–E, 37

Copelatus politus Sharp, 1882: 568.

Copelatus politus Sharp, 1882: Régimbart 1899: 292-293 (descr.); Zimmermann 1919: 198 (cat.), 1920: 145 (cat.); Guéorguiev 1968: 34 (cat.), 1978: 269 (key); Nilsson 2001: 66 (cat.).

Exocelina polita (Sharp, 1882): Nilsson and Fery 2006: 56 (n. comb.).

Type locality.

Indonesia: West Papua Province: Manokwari Regency, Arfak Mts., Hatam. Note: Hatam is situated in the Arfak Mountains north of Ransiki (A. Riedel, pers. comment).

Type material.

Lectotype (hereby designated): male “Type” [round, with red rim], "Hatam N. Guinea Luglio [= July] 1875. Beccari." [hw, Beccari], "Sharp Coll 1905-313", "Hatam, New-Guinea July 1875 Beccari 660" [hw, Sharp], "Type 660 Copelatus politus n. sp. New Guinea" [hw, Sharp], "Lectotype Copelatus politus Sharp des. Shaverdo, Hendrich & Balke 2012" [red, printed] (BMNH). Notes: The lectotype is designated in order to support the stability of nomenclature since it is not clear from the original description that it has been based on the single male. The genialia are partly damaged.

Diagnosis.

Beetle middle-sized, brown, shiny, with almost invisible dorsal punctation; pronotum with distinct lateral bead; male antennomeres 3-4 strongly enlarged and triangular (3 distinctly larger than 4), 5-6 distinctly enlarged, 7 somewhat enlarged; male protarsomere 4 with small, thin, slightly curved anterolateral hook; median lobe with strong submedian constriction in ventral view, apex of median lobe elongate in lateral view and broader in ventral view; paramere with notch on dorsal side and subdistal part short and small, with not numerous, relatively short, thick, and flattened setae.

Redescription.

Size and shape: Beetle middle-sized (TL-H 3.9 mm, TL 4.35 mm, MW 2.15 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Dorsally dark brown, with reddish head, pronotal sides, and sutural bands on elytra; head appendages yellowish red, legs darker, especially hind legs (Fig. 37).

Surface sculpture: Head with dense punctation (spaces between punctures 1-2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller or equal than diameter of cells of microreticulation. Pronotum with distinctly finer, sparser, and more evenly distributed punctation than on head. Elytra with very sparse and fine punctation, almost invisible. Head, pronotum, and elytra with strongly impressed microreticulation, dorsal surface shiny. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal sternites with distinct microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal sternites.

Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded and smooth anteriorly, with very small anterolateral extensions. Blade of prosternal process lanceolate, relatively broad, convex, with distinct bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal sternite 7 broadly rounded apically.

Male: Antennomeres 3-4 strongly enlarged and triangular (3 distinctly larger than 4), 5-6 distinctly enlarged, 7 somewhat enlarged (Fig. 11A); antennomeres 3-7 rugose ventrally. Protarsomere 4 with small, thin, slightly curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 12 short setae and posterior row 5 short setae (Fig. 11B). Abdominal sternite 7 with 3-4 lateral striae on each side. Median lobe with strong submedian constriction in ventral view, apex of median lobe elongate in lateral view and broader in ventral view (Figs 11C, D). Paramere with notch on dorsal side and subdistal part short and small, with not numerous, relatively short, thick, and flattened setae (Fig. 11E).

Female: Unknown.

Distribution.

Indonesia: West Papua Province: Manokwari Regency. The species is known only from the lectotype from the Arfak Mountains, the eastern part of Bird’s head (Fig. 50).