Cryptoerithus occultus Rainbow

Platnick, N. I. & Baehr, B., 2006, A Revision Of The Australasian Ground Spiders Of The Family Prodidomidae (Araneae: Gnaphosoidea), Bulletin of the American Museum of Natural History 2006 (298), pp. 1-287 : 235-248

Cryptoerithus occultus Rainbow Figures 22, 553–557; Map 39

Cryptoerithus occultus Rainbow, 1915: 778 , pl. 67, figs. 3–5 (female holotype from Flat Rockhole, South Australia, in SAM, examined).

DIAGNOSIS: This may be the western sister species of C. lawlessi ; males differ in having a longer, narrower median apophysis (fig. 554), females in lacking pronounced posterior epigynal margins (fig. 556) and having larger anterior epigynal ducts (fig. 557).

MALE: Total length 3.16. Carapace 1.26 long, 1.06 wide, 0.40 high, length/width 1.18; sternum 0.90 long, 0.68 wide, length/ width 1.32; abdomen 1.90 long, 0.90 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.85:0.76:1.12. Body, legs pale yellow. Eye group width 0.77 of head width; AME 0.12; ALE 0.12; PME 0.16; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.01; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.32; AME–AME 0.26; PME–PME 0.30. Clypeus 0.01 high. Cheliceral promargin, retromargin each with one tiny tooth. Abdomen weakly covered with gray plumose setae, venter with unordered, stiff, erect setae; ALS 0.22 of abdominal length, about half their diameter apart. Tarsi I, II proventrally with band of scaled setae. Palp (figs. 553–555): cymbium retrolaterally straight with basal hook; conductor originating prolaterally, long, semicircular; median apophysis long, thin, with basal hook, slender, bent tip; terminal apophysis conical, with sharp tip, basally situated, connected to embolar base; sperm duct semicircular; embolus thin, semicircular, separated from tegulum; retrolateral tibial apophysis rectangular.

FEMALE: Total length 4.74. Carapace 1.78 long, 1.48 wide, 0.54 high, length/width 1.20; sternum 1.20 long, 0.96 wide, length/ width 1.25; abdomen 2.96 long, 1.56 wide; coxa I 0.66 long; relative length of coxae I–IV 1.00:0.90:0.87:1.30. Coloration as in male. Eye group width 0.62 of head width; AME 0.14; ALE 0.14; AME–AME 0.04; PME–PME 0.02; ALE–PLE 0.06; AME– AME 0.30. Clypeus 0.01 high. Chelicerae with two small promarginal, three strong retromarginal teeth. ALS 0.24 of abdominal length. Epigynum (figs. 556, 557) with narrow epigynal hood; lateral plates rectangular, contiguous, spermathecae inverted v-shaped.

MATERIAL EXAMINED: Northern Territory: 29 km ESE Uluru, Uluru-Kata Tjuta National Park, 25 ° 219S, 131 ° 229E, pitfall, Oct. 1994 (Horn Expedition Centenary Survey, NMV K8838 , K8839 ), 1 Oi, 1♀. South Australia: 10 km N Anvil Hole Native Well, Witjira National Park , 26 ° 169S, 135 ° 419E, Nov. 19–25, 1995, pitfall ( SAM N1997 /156), 1 Oi ; 12.2 km NW Cheesman Peak , 27 ° 209S, 130 ° 149E, Oct. 22–25, 1996, pitfall, sand plain ( SAM NN10962 ), 1 Oi ; Flat Rockhole, Musgrave Ranges , ca. 26 ° 359S, 132 ° 309E, July 13, 1914 (S. White, SAM N1981 /376), 1♀ (holotype) ; 2.7 km SSE Invasion Tank , 26 ° 069S, 140 ° 509E, Nov. 9, 1994 ( SAM NN11774 ), 1♀ ; 6 km N Mirra Mitta Bore, Cowarie Station , 27 ° 439S, 138 ° 489E, Apr. 23– 29, 1995, pitfall ( SAM N1997 /158), 1♀ ; 6 km W Moomba , 28 ° 079S, 140 ° 099E, Nov. 24, 1999 ( SAM NN11794–11796 ), 2 Oi, 1♀ ; 8 km E Mount Finke , near salt lake, 30 ° 559S, 134 ° 029E, Oct. 2, 1988, pitfall (D. Hirst, SAM N1997 /157), 1 Oi ; 31 km NW Renmark, 33 ° 599S, 140 ° 309E, May 2–June 8, 1995, flight intercept trap (K. Pullen, ANIC), 1 Oi ; 1 km W Vokes Hill Corner, Great Victoria Desert , 28 ° 349S, 130 ° 419E, Apr. 14–19, 1994 (D. Hirst, J. Forrest, SAM NN11792 ), 1 Oi ; 11.5 km SE Wares Peak, 29 ° 409S, 135 ° 469E, Sept. 29–Oct. 5, 1995, pitfall (H. Owens, SAM NN11770 , 11772 ), 2♀. Western Australia: Barlee Range Nature Reserve , 23 ° 069S, 115 ° 459E, Aug. 1993, pitfalls (S. van Leeuwen, B. Bromilow, WAM T45261), 2 Oi ; Barrow Island, John Wayne Country , 20 ° 459S, 115 ° 229E, Nov. 4–30, 1993, sandy site, pitfall (M. Harvey, J. Waldock, WAM T45128), 1♀ ; Bidgemia Station, Gasgoyne Junction , 25 ° 039S, 115 ° 189E, Aug. 17– Oct. 5, 1994, pitfall (A. Sampey, WAM T44952), 1♀ ; Bidgemia Station, Gasgoyne Junction , 25 ° 059S, 115 ° 229E, Aug. 17–Oct. 5, 1994, pitfall (A. Sampey, WAM T44951), 1 Oi ; Bidgemia Station, Gasgoyne Junction , 25 ° 079S, 115 ° 269E, Aug. 17–Oct. 6, 1994, pitfalls (A. Sampey, WAM T44950), 2 Oi, 3♀ ; Boolathana Station , 24 ° 259S, 113 ° 409E, Sept. 30, 1994 – Jan. 15, 1995, pitfall (N. McKenzie, J. Rolfe, WAM 99 /654–656), 3 Oi ; Boolathana Station , 24 ° 259S, 113 ° 419E, Sept. 30, 1994 – Jan. 15, 1995, pitfalls (N. McKenzie, J. Rolfe, WAM T44953, 44954 ), 5 Oi, 1♀ ; Boolathana Station , 24 ° 259S, 113 ° 429E, Aug. 20–Sept. 30, 1994, pitfall (A. Sampey, WAM T44955), 1 Oi ; Boolathana Station , 24 ° 259S, 113 ° 459E, Sept. 30, 1994 – Jan. 15, 1995, pitfalls (N. McKenzie, J. Rolfe, WAM T44956), 2 Oi, 1♀ ; Bunline Rocks Nature Reserve , 29 ° 599S, 116 ° 369E, May 22–Sept. 17, 1996, pitfall (M. Harvey, J. Waldock, WAM T45138), 1♀ ; Bush Bay , 25 ° 089S, 113 ° 499E, Aug. 16–Sept. 29, 1994, pitfall (M. Harvey, WAM T44957), 1♀, May 23–Aug. 23, 1995, pitfall (N. Hall, WAM T44958), 1♀ ; Durokoppin Nature Reserve , 31 ° 309S, 117 ° 449E, Nov. 3–14, 1988 (D. Mitchell, WAM 99 /667), 1 Oi ; Francois Peron National Park, 25 ° 499S, 113 ° 329E, Aug. 24–Oct. 11, 1994, pitfall (A. Sampey, WAM T44964), 1 Oi , Oct. 11, 1994 – Jan. 18, 1995 (N. McKenzie, J. Rolfe, WAM T44965), 1♀ ; Francois Peron National Park , 25 ° 539S, 113 ° 339E, Jan. 17–May 25, 1995, pitfall (M. Harvey, WAM T44963), 1 Oi ; Francois Peron National Park, 25 ° 599S, 113 ° 349E, Aug. 24–Oct. 10, 1994, pitfall (A. Sampey, WAM T44967), 1♀, Jan. 18–May 26, 1995, pitfalls (M. Harvey, WAM T44966, 45573 ), 3 Oi , 1♀, May 26–Aug. 30, 1995, pitfall (N. Hall, WAM T44968), 1♀ ; Kennedy Range National Park, 24 ° 309S, 115 ° 019E, Oct. 6, 1994 – Jan. 14, 1995, pitfall (N. McKenzie, J. Rolfe, WAM T44970), 1 Oi , Jan. 14–May 29, 1995, pitfall (P. West, WAM T44971), 1♀ ; Kennedy Range National Park , 24 ° 309S, 115 ° 029E, May 29–Aug. 28, 1995, pitfall (N. Hall, WAM T44969), 1♀ ; Kennedy Range National Park , 24 ° 339S, 114 ° 589E, Aug. 18–Oct. 4, 1994, pitfalls (M. Harvey, WAM T44972), 3 Oi ; 39 km E Laverton , 28 ° 289S, 122 ° 509E, Nov. 2–3, 1990, pitfall (E. Pianka, WAM 91 / 1043), 1♀ ; Little Sandy Desert , 17.5 km SE Burranbar Pool, 23 ° 559S, 120 ° 329E, Apr. 1997, pitfalls (S. van Leeuwen, B. Bromilow, WAM T45626), 1 Oi, 1♀ ; Mardathuna Station , 24 ° 249S, 114 ° 279E, Aug. 19–Oct. 5, 1994, pitfall (P. West, WAM 99 /661–664), 4 Oi ; Mardathuna Station, 24 ° 249S, 114 ° 289E, Jan. 14– May 24, 1995, pitfalls (A. Sampey, WAM T44975, 44976 ), 4 Oi , May 24–Aug. 26, 1995, pitfall (N. Hall, WAM T44977), 1 Oi ; Mardathuna Station , 24 ° 269S, 114 ° 309E, Oct. 5, 1994 – Jan. 14, 1995, pitfall (N. McKenzie, J. Rolfe, WAM T44974), 1♀ ; Mardathuna Station, 24 ° 279S, 114 ° 319E, Aug. 19–Oct. 5, 1994, pitfall (P. West, WAM 99 /657, 658), 2 Oi , Oct. 5, 1994 – Jan. 14, 1995, pitfall (N. McKenzie, J. Rolfe, WAM 99 /660), 1 Oi ; Mardathuna Station , 24 ° 319S, 114 ° 389E, Aug. 19– Oct. 5, 1994, pitfall (P. West, WAM T44973), 1 Oi ; Meedo Station , 25 ° 419S, 114 ° 379E, Aug. 22–Oct. 11, 1994, pitfalls (P. West, WAM T44978), 1 Oi, 1♀ ; Nanga Station , 26 ° 299S, 114 ° 039E, Jan. 19–May 11, 1995, pitfall (A. Sampey, WAM T44979), 1♀ ; Nerren Nerren Station , 27 ° 009S, 114 ° 329E, Oct. 15, 1994 – Jan. 11, 1995, pitfall (N. McKenzie, J. Rolfe, WAM T44980), 1♀ ; Quobba Station, Cape Cuvier, 24 ° 129S, 113 ° 279E, Aug. 21–Sept. 29, 1994, pitfall (P. West, WAM T44962), 1 Oi , Jan. 15–May 29, 1995, pitfall (A. Sampey, WAM 99 /653, T44961), 2 Oi ; Quobba Station, Cape Cuvier , 24 ° 139S, 113 ° 309E, Aug. 21– Sept. 29, 1994, pitfall (P. West, WAM T44960), 1 Oi ; Quobba Station, Cape Cuvier , 24 ° 159S, 113 ° 339E, Aug. 21–Sept. 29, 1994, pitfall (P. West, WAM T44959), 1 Oi ; Woodleigh Station , 26 ° 119S, 114 ° 319E, Jan. 12–May 17, 1995, pitfall (P. West, WAM T44913), 1 Oi ; Woodleigh Station , 26 ° 129S, 114 ° 259E, Aug. 22–Oct. 10, 1994, pitfall (M. Harvey, WAM T44984), 1 Oi ; Woodleigh Station, 26 ° 139S, 114 ° 359E, Aug. 22–Oct. 12, 1994, pitfall (M. Harvey, WAM T44982), 1 Oi , May 17–Aug. 21, 1995, pitfall (N. Hall, WAM T44983), 1♀ ; Woodleigh Station , 26 ° 139S, 114 ° 369E, Aug. 22–Oct. 11, 1994, pitfall (M. Harvey, WAM T44981), 1 Oi .

DISTRIBUTION: Widespread across the southwest two-thirds of Australia (map 39).

Cryptoerithus nichtaut , new species

Figures 558–562; Map 39

TYPE: Female holotype taken in pitfall trap in a dry eucalypt woodland at Thatch Creek , 19 ° 069S, 145 ° 189E, Queensland (July 26–Dec. 1, 1992; R. Raven, P., E. Lawless, M. Shaw), deposited in QMB ( S25300 ) .

ETYMOLOGY: The specific name is an arbitrary combination of letters.

DIAGNOSIS: Males and females have not been collected together, but are matched on the basis of their highly hirsute anterior legs. Males have a very long, distally sinuous embolus (fig. 559) and a short, wide, triangular retrolateral tibial apophysis (fig. 560), females have a rectangular epigynum (fig. 561).

MALE: Total length 2.96. Carapace 1.32 long, 1.16 wide, 0.46 high, length/width 1.13; sternum 0.90 long, 0.74 wide, length/width 1.21; abdomen 1.64 long, 1.02 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.84:0.72:1.20. Cephalothorax, legs pale; abdomen grayish, with weak orange brown scutum, venter pale. AME elevated; eye group width 0.80 of head width; AME 0.16; ALE 0.12; PME 0.18; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.00; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.36; AME–AME 0.34; PME– PME 0.32. Clypeus 0.08 high. Abdomen covered with gray plumose setae, ventrally with two longitudinal rows of stiff, erect setae; ALS 0.27 of abdominal length, about half their diameter apart. Tibiae, metatarsi, tarsi I, II proventrally with long dense scopula. Palp (figs. 558–560): cymbium with dorsoapical scopula; conductor originating prolaterally, long, s-shaped; median apophysis elongate, about four times longer than wide, with small distal hook; terminal apophysis absent; sperm duct strongly s-shaped; embolus thin, semicircular, tip s-shaped, originating retrolaterally, embolar base separated from tegulum; retrolateral tibial apophysis short, broad, triangular, with ventrally bent tip.

FEMALE: Total length 3.74. Carapace 1.50 long, 1.36 wide, 0.42 high, length/width 1.10; sternum 1.06 long, 0.80 wide, length/width 1.32; abdomen 2.24 long, 1.44 wide; coxa I 0.54 long; relative length of coxae I–IV 1.00:0.78:0.74:1.22. Coloration as in male. Eye group width 0.73 of head width; PME–PLE 0.06; ALE–PLE 0.04; eye group AME–PME 0.38. ALS 0.3 of abdominal length. Epigynum (figs. 561, 562): atrium broad, slit-like, anterior margin widely m-shaped, with narrow epigynal hood, lateral plates rectangular, contiguous; spermathecae inverted v-shaped.

OTHER MATERIAL EXAMINED: Queensland: Danbulla State Forest , 17 ° 129S, 145 ° 409E, Aug. 6, 1992 (J. Murphy, JAM), 1♀ ; Jack’s Hill Gorge, Broken River , 19 ° 289S, 144 ° 469E, July 21–Sept. 20, 1995, pitfall, vine thicket on limestone (P. Lawless, A. Cook, C. McHenry, QMB S40889 ), 1Oi ; Mazeppa National Park , SW corner near road, 22 ° 169S, 147 ° 169E, May 11, 2000 (G. Milledge, H. Smith, AMS KS57817 ), 1Oi .

DISTRIBUTION: Known only from north- and mideastern Queensland (map 39).

Cryptoerithus hasenpuschi , new species

Figures 563–567; Map 40

TYPE: Male holotype taken in pitfall trap in Forty Mile Scrub National Park , 18 ° 059S, 144 ° 519E, Queensland (Oct. 1993 – Jan. 1994; J. Hasenpusch), deposited in QMB ( S52087 ) .

ETYMOLOGY: The specific name is a patronym in honor of Jack Hasenpusch, the collector of the type.

DIAGNOSIS: Males differ from those of C. nichtaut and C. lawlessi is having the embolus situated closely against the other palpal elements throughout its length (fig. 564); females differ from those of C. nichtaut in the more triangular anterior elements of the internal genitalia (fig. 567).

MALE: Total length 2.84. Carapace 1.30 long, 1.06 wide, 0.30 high, length/width 1.22; sternum 0.92 long, 0.70 wide, length/ width 1.31; abdomen 1.54 long, 1.02 wide; coxa I 0.46 long; relative length of coxae I–IV 1.00:0.87:0.78:1.22. Carapace cinnamon brown, with dark filigree net pattern; sternum, chelicerae, endites, labium pale; abdomen dorsally gray, with weak orange brown scutum, venter pale; legs cinnamon brown, mottled with gray. Eye group width 0.86 of head width; AME 0.12; ALE 0.12; PME 0.18; PLE 0.08; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.00; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.30; AME–AME 0.26; PME–PME 0.30. Clypeus 0.02 high. Abdomen dorsally covered with gray plumose setae, venter with two longitudinal rows of stiff, erect setae; ALS 0.35 of abdominal length, contiguous. Tarsi I, II proventrally with band of short, bent setae. Palp (figs. 563–565): cymbium retrolaterally straight, with basal hook, dorsoapical scopula; conductor originating prolaterally, long, semicircular; median apophysis cane-shaped, about twice as long as wide, with tiny hook; terminal apophysis prolaterally situated, triangular, with slender, sharp tip; sperm duct bifurcate; embolus thin, semicircular, originating prolaterally; retrolateral tibial apophysis short, broad, triangular, with ventrally bent tip.

FEMALE: Total length 3.38. Carapace 1.40 long, 1.16 wide, 0.36 high, length/width 1.21; sternum 0.92 long, 0.80 wide, length/ width 1.15; abdomen 1.98 long, 1.10 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.80:0.76:1.20. Body, legs pale. Eye group width 0.55 of caput width; AME 0.10; ALE 0.10; PME 0.14; PLE 0.08; AME–AME 0.02; AME–ALE 0.02; PME– PME 0.00; PME–PLE 0.02; ALE–PLE 0.02; eye group AME–PME 0.32; AME–AME 0.22; PME–PME 0.20. Clypeus 0.02 high. Abdomen covered with gray plumose setae, frontally with band of long, bent spines, venter with two longitudinal rows of stiff, erect setae; ALS 0.35 of abdominal length, contiguous. Tarsi I, II proventrally with band of short, bent setae. Epigynum (figs. 566, 567) with narrow anterior epigynal hood, rectangular, contiguous lateral plates; epigynal ducts short, spermathecae separated by more than their diameter, sausage-shaped, twisted, in inverted v-shaped position.

OTHER MATERIAL EXAMINED: Queensland: 50 km W Mount Garnet , 17 ° 409S, 145 ° 079E, Apr. 14–July 19, 2002, pitfall (J. Hasenpusch, QMB S67694 ), 1Oi ; Cape Cleveland Road , 19 ° 219S, 147 ° 019E, July 27–Dec. 2, 1992 (R. Raven, P., E. Lawless, M. Shaw, QMB S21709 ), 1♀ .

DISTRIBUTION: Known only from northeastern Queensland (map 40).

Cryptoerithus quamby , new species

Figures 568–572; Map 39

TYPE: Female holotype taken in grazed site with remnant eucalypt, Acacia , Leptospermum and some open grassy areas at Dugald River crossing, N Quamby, 20 ° 129S, 140 ° 149E, Queensland (July 12, 1995; J. Thompson, M. Tio, S. Cowan), deposited in AMS (KS44181).

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Females resemble those of C. nichtaut in having highly hirsute anterior legs, but have a posterior ledge on the epigynum (fig. 571) and much smaller anterior internal epigynal elements (fig. 572). Males resemble those of C. nonaut but have a longer embolus, originating retrolaterally rather than proximally (figs. 569, 570).

MALE: Total length 2.82. Carapace 1.40 long, 1.20 wide, 0.32 high, length/width 1.16; sternum 0.92 long, 0.72 wide, length/width 1.27; abdomen 1.42 long, 1.12 wide; coxa I 0.54 long; relative length of coxae I–IV 1.00:0.77:0.66:1.11. Carapace pale yellow, with marginal dark filigree net pattern; sternum yellow; chelicerae gray; endites, labium gray, distally pale; abdomen gray, dorsally with weak orange brown scutum, half moon-shaped pale spot in front of spinnerets, venter pale; legs pale, mottled with gray on distal half of femora. Eye group width 0.63 of head width; AME 0.14; ALE 0.14; PME 0.20; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.00; PME– PLE 0.04; ALE–PLE 0.02; eye group AME– PME 0.36; AME–AME 0.30; PME–PME 0.36. Clypeus 0.08 high. Abdomen covered with gray plumose setae; ALS 0.35 of abdominal length, about half their diameter apart. Metatarsi, tarsi I, II with scopula. Palp (figs. 568–570): cymbium with dorsoapical scopula; conductor originating prolaterally, long, straight, with sharp tip; median apophysis about twice as long as wide, with bent, slender tip; terminal apophysis conical, medially situated, connected to embolar base; sperm duct strongly s-shaped; embolus thin, semicircular, originating basally, embolar base separated from tegulum; retrolateral tibial apophysis triangular, with deep dorsal concavity, additional basal apophysis.

FEMALE: Total length 3.92. Carapace 1.56 long, 1.48 wide, 0.50 high, length/width 1.05; sternum 1.02 long, 0.82 wide, length/ width 1.24; abdomen 2.36 long, 1.34 wide; coxa I 0.62 long; relative length of coxae I–IV 1.00:0.90:0.80:1.16. Carapace pale yellow, with marginal dark filigree net pattern;

sternum yellow; chelicerae gray; endites, labium gray, distally pale; abdomen gray, venter pale; legs pale, mottled with gray on distal half of femora. Eye group width 0.63 of caput width; AME 0.12; ALE 0.12; PME 0.20; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.01; PME– PLE 0.04; ALE–PLE 0.04; eye group AME– PME 0.38; AME–AME 0.26; PME–PME 0.34. Clypeus 0.08 high. Abdomen covered with gray plumose setae, frontally with band of long, bent spines, venter with two longitudinal rows of stiff, erect setae; ALS 0.36 of abdominal length, about half their diameter apart. Tibiae, metatarsi, tarsi I, II proventrally with long dense scopula. Epigynum (figs. 571, 572): atrium broad, slitlike, with broad posterior ledge; epigynal ducts thin, irregularly curved, spermathecae sausage-shaped, twisted, in horizontal position.

OTHER MATERIAL EXAMINED: Queensland: Meta Park, 23 ° 449S, 146 ° 549E, Oct. 20, 1998 (QMB S67725 ), 1Oi.

DISTRIBUTION: Known only from northwestern and central Queensland (map 39).

Cryptoerithus lawlessi , new species

Figures 573–577; Map 40

TYPE: Female holotype taken in pitfall trap in vine thicket on hill in Taroom District , 25 ° 259S, 149 ° 589E, Queensland (Sept. 10–Nov. 12, 1996; P. Lawless), deposited in QMB ( S61303 ) .

ETYMOLOGY: The specific name is a patronym in honor of the collector of the type and many other interesting prodidomids.

DIAGNOSIS: Males differ from those of C. nichtaut in having a shorter, more retrolaterally situated median apophysis and much longer tegular apophysis (fig. 574), and by the angular retrolateral tibial apophysis (fig. 575); females resemble those of C. cleveland in lacking highly hirsute anterior legs, but have posteriorly much more pronounced epigynal plates (fig. 576) and more anterior internal epigynal elements (fig. 577).

MALE: Total length 2.70. Carapace 1.20 long, 1.04 wide, 0.34 high, length/width 1.15; sternum 0.90 long, 0.68 wide, length/width 1.32; abdomen 1.50 long, 0.84 wide; coxa I 0.44 long; relative length of coxae I–IV 1.00:0.86:0.68:1.18. Carapace pale yellow, with dark filigree net pattern; sternum, mouthparts pale, abdomen gray, venter pale, legs mottled with gray. Eye group width 0.72 of head width; AME 0.10; ALE 0.10; PME 0.18; PLE 0.08; AME–AME 0.02; AME– ALE 0.02; PME–PME 0.00; PME–PLE 0.02; ALE–PLE 0.02; eye group AME–PME 0.36; AME–AME 0.22; PME–PME 0.28. Clypeus 0.02 high. Abdomen covered with gray plumose setae, ventrally with two longitudinal rows of stiff, erect setae; ALS 0.48 of abdominal length, contiguous. Metatarsi, tarsi I, II proventrally with band of short bent setae. Palp (figs. 573–575): cymbium retrolaterally straight, with basal hook, dorsoapical scopula; conductor originating distally, broadly ellipsoid; median apophysis cane-shaped, about twice as long as wide; terminal apophysis triangular, with sharp tip, prolaterally situated; sperm duct weakly sshaped; embolus thin, semicircular, originating prolaterally; retrolateral tibial apophysis short, broad, triangular, with ventrally bent tip.

FEMALE: Total length 3.32. Carapace 1.40 long, 1.26 wide, 0.34 high, length/width 1.11; sternum 1.00 long, 0.80 wide, length/ width 1.25; abdomen 1.92 long, 1.04 wide; coxa I 0.52 long; relative length of coxae I–IV 1.00:0.88:0.77:1.15. Coloration as in male. Eye group width 0.68 of head width; PME– PME 0.01; PME–PLE 0.04; eye group AME– PME 0.2. ALS 0.36 of abdominal length. Epigynum (figs. 576, 577) with narrow epigynal hood, lateral plates inverted ushaped; spermathecae widely separated.

OTHER MATERIAL EXAMINED: Queensland: Agnes Waters , 24 ° 139S, 151 ° 559E, July 1992 (J. Wunderlich, QMB S21214 ), 1 Oi, 1♀ ; Bushley Station , 23 ° 329S, 150 ° 159E, July 19–Oct. 22, 1990, open forest pitfall (D. Wallace, R. Raven, K. Williams, QMB S25616 ), 1 Oi ; Newcastle Range, E Georgetown , 18 ° 349S, 143 ° 519E, Feb. 2–June 29, 2002, pitfall (J. Hasenpusch, QMB S62435), 1♀ ; Yarmina , 21 ° 499S, 146 ° 299E, Mar. 1, 1999, tree clearing (T. Churchill, QMB S67726 ), 1♀ .

DISTRIBUTION: Apparently widespread in Queensland (map 40).

Cryptoerithus griffith , new species

Figures 578, 579; Map 40

TYPE: Female holotype taken at Griffith University Study Area, Brisbane , 27 ° 289S, 153 ° 019E, Queensland (Dec. 11, 1976; G. Oliver), deposited in QMB ( S61304 ) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Females of this distinctive species can easily be recognized by the distally darkened femora, patellae, and tibiae on the highly hirsute anterior legs, as well as by the wide anterior epigynal margin, the large, inverted u-shaped posterior projections, and the bulb-shaped anterior internal epigynal elements (figs. 578, 579).

MALE: Unknown.

FEMALE: Total length 3.26. Carapace 1.40 long, 1.21 wide, 0.40 high, length/width 1.13; sternum 1.00 long, 0.76 wide, length/ width 1.31; abdomen 1.86 long, 1.34 wide; coxa I 0.56 long; relative length of coxae I–IV 1.00:0.75:0.71:1.14. Carapace cinnamon brown, with dark margin, dark chevrons on top; sternum orange, lateral margins darker; chelicerae, endites, labium gray, distally pale; abdomen gray, with small pale spot in front of spinnerets; legs cinnamon brown with

distally darkened femora, patellae, tibiae. Eye group width 0.77 of head width; AME 0.12; ALE 0.12; PME 0.18; PLE 0.10; AME– AME 0.06; AME–ALE 0.02; PME–PME 0.00; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.36; AME–AME 0.30; PME–PME 0.32. Clypeus 0.04 high. Abdomen covered with gray plumose setae; ALS 0.32 of abdominal length, about half their diameter apart. Tibia I distal half, metatarsi, tarsi I, II proventrally with scopula. Epigynum (figs. 578, 579): anterior margin with inverted v-shaped epigynal hood, posterior margin with one pair of large anteriorly rounded projections; epigynal ducts and spermathecae in x-shaped position.

OTHER MATERIAL EXAMINED: South Australia: Billiatt Conservation Park, 34 ° 599S, 140 ° 289E, Nov. 19, 1996, vibration (D. Hirst, SAM NN11782), 1♀.

DISTRIBUTION: Known only from southeastern Queensland and eastern South Australia (map 40).

Cryptoerithus shadabi , new species

Figures 580–584; Map 41

TYPE: Male holotype taken in drift fence pitfall trap at site 9.2 km SSE of the confluence of Lindsay River and Mullaroo Creek , 34 ° 129S, 141 ° 109E, Victoria (Nov. 1985; A. Yen), deposited in NMV (Ent. 362) .

ETYMOLOGY: The specific name is a patronym in honor of Dr. Mohammad U. Shadab, who supplied the drawings for this monograph.

DIAGNOSIS: Males and females have not been collected together and are only tentatively matched here. Males have a bifid tegular apophysis and a greatly shortened embolus (fig. 581); females have a y-shaped epigynal atrium (fig. 583) and widely separated spermathecae (fig. 584).

MALE: Total length 3.72. Carapace 1.50 long, 1.26 wide, 0.30 high, length/width 1.19; sternum 1.00 long, 0.82 wide, length/ width 1.22; abdomen 2.22 long, 1.22 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.96:0.8:1.36. Carapace cinnamon brown, with dark filigree net pattern; sternum, mouthparts pale; abdomen gray, venter pale; legs cinnamon brown. Eye group width 0.67 of head width; AME 0.10; ALE 0.12; PME 0.16; PLE 0.10; AME–AME 0.06; AME–ALE 0.02; PME–PME 0.00; PME–PLE 0.04; ALE–PLE 0.04; eye group AME–PME 0.32; AME–AME 0.26; PME–PME 0.28. Clypeus 0.04 high. Cheliceral margins without teeth; labium triangular. Abdomen dorsally covered with gray plumose setae, ventrally with two longitudinal rows of stiff, erect setae; ALS 0.32 of abdominal length, contiguous. Metatarsi, tarsi I, II proventrally with band of short bent setae. Palp (figs. 580–582): cymbium retrolaterally straight, with basal hook, dorsoapical scopula; conductor originating medially, broadly ellipsoid, distal part concave; median apophysis cane-shaped, about twice as long as wide; terminal apophysis bipartite, prolaterally situated; sperm duct semicircular; embolus finger-shaped, with sharp tip, originating prolaterally, embolar base hidden behind terminal apophysis; retrolateral tibial apophysis short, broad, triangular, with ventrally bent tip.

FEMALE: Total length 4.00. Carapace 1.64 long, 1.44 wide, 0.32 high, length/width 1.13; sternum 1.20 long, 0.86 wide, length/ width 1.39; abdomen 2.36 long, 1.24 wide; coxa I 0.64 long; relative length of coxae I–IV 1.00:0.81:0.78:1.25. Coloration as in male. Eye group width 0.61 of head width; AME 0.11; PME 0.18. Clypeus 0.02 high. ALS 0.42 of abdominal length. Epigynum (figs. 583, 584): atrium y-shaped, formed by two lateral plates, with narrow epigynal hood; spermathecae widely separated.

OTHER MATERIAL EXAMINED: South Australia: Billiatt Conservation Park , 34 ° 589S, 140 ° 279E, Nov. 19, 1996, vibration, mallee litter (D. Hirst, J. Forrest, SAM N1997 /160, NN11819, 11820), 3♀ ; Lake Gilles Conservation Park , 33 ° 059S, 136 ° 399E, May 21, 1995, vibration (D. Hirst, SAM NN11790 ), 1♀ ; 4 km N Marble Range Homestead , 34 ° 269S, 135 ° 319E, Mar. 29, 1987 (D. Lee, D. Hirst, SAM N1989 / 114), 1 Oi. Western Australia: Sieda, E Grass Patch, 33 ° 149S, 121 ° 469E, Dec. 1, 1996 – Feb. 28, 1997, pitfall, water reserve (A. Longbottom, WAM T45102), 1♀ .

DISTRIBUTION: Widespread across southern Australia (map 41).

Cryptoerithus stuart , new species

Figures 585–589; Map 41

TYPE: Female holotype taken under log along Stuart Highway , 16 ° 599S, 133 ° 269E, Northern Territory (May 19, 1999; M. Gray, G. Milledge, H. Smith), deposited in AMS ( KS57272 ) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: The two sexes have not been collected together and are only tentatively matched here; their differences in leg scopulae may signal a mismatch. Males have a distinctively narrow, sinuous retrolateral tibial apophysis (fig. 587); females resemble those of C. lawlessi but have a much smaller epigynal hood and a much wider median epigynal septum (fig. 588).

MALE: Total length 2.30. Carapace 1.08 long, 0.96 wide, 0.48 high, length/width 1.12; sternum 0.76 long, 0.60 wide, length/width 1.27; abdomen 1.22 long, 0.76 wide; coxa I 0.36 long; relative length of coxae I–IV 1.00:0.83:0.72:1.28. Carapace pale, with marginal dark filigree net pattern; sternum, endites, labium pale; chelicerae gray; abdomen gray, venter pale; legs pale, mottled with gray. Eye group width 0.74 of head width; AME 0.11; ALE 0.10; PME 0.18; PLE 0.08; AME– AME 0.10; AME–ALE 0.02; PME–PME 0.00; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.28; AME–AME 0.24; PME–PME 0.28. Clypeus 0.06 high. Cheliceral promargin, retromargin each with three tiny teeth; labium triangular. Abdomen dorsally covered with gray plumose setae, ventrally with two longitudinal rows of stiff, erect setae; ALS 0.33 of abdominal length, about their diameter apart. Metatarsi, tarsi I, II with scopula. Palp (figs. 585–587): cymbium with dorsoapical scopula; conductor originating prolaterally, long, semicircular, with sharp tip; median apophysis cane-shaped, about twice as long as wide; terminal apophysis medially situated, triangular, with slender, sharp tip; sperm duct weakly s-shaped; embolus thin, semicircular, originating basally; retrolateral tibial apophysis slender, weakly sshaped; femur without ventral process.

FEMALE: Total length 2.40. Carapace 1.26 long, 1.16 wide, 0.50 high, length/width 1.08; sternum 0.88 long, 0.74 wide, length/ width 1.19; abdomen 1.14 long, 0.64 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.8:0.72:1.00. Carapace pale yellow, with marginal dark filigree net pattern; sternum, endites, labium pale yellow; chelicerae gray; abdomen gray, venter pale; legs pale, mottled with gray. Eye group width 0.77 of caput width; AME 0.14; ALE 0.12; PME 0.20; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.01; PME–PLE 0.04; ALE– PLE 0.02; eye group AME–PME 0.34; AME–AME 0.30; PME–PME 0.34. Clypeus 0.08 high. Cheliceral promargin with five, retromargin with three tiny teeth; labium triangular. Abdomen covered with gray or plumose setae, frontally with band of long, bent spines, venter with two longitudinal rows of stiff, erect setae; ALS 0.49 of abdominal length, about their diameter apart. Tibia I distal half, metatarsi, tarsi I, II proventrally with scopula. Epigynum (figs. 588, 589) with narrow anterior hood, posterior margin with elevated median septum, rectangular lateral plates; epigynal ducts flat, helical, spermathecae widely separated, more than their diameter apart, sausageshaped, twisted, in horizontal position.

OTHER MATERIAL EXAMINED: Northern Territory: Kidman Springs Station , 16 ° 079S, 130 ° 579E, May 1997, red loam soil (T. Churchill, MNT A001566 ), 1Oi, mid Apr. 1998, pitfall (T. Churchill, MNT A001523 ), 1Oi ; Mitchell Grasslands, Mount Sanford, Barkly Tablelands , 16 ° 599S, 130 ° 329E, May 1997, pitfall (A. Fisher, MNT A001550 ), 2Oi ; Willeroo , 15 ° 179S, 131 ° 359E, July 3–9, 1996, pitfall, clay (T. Churchill, MNT A002958 ), 1Oi .

DISTRIBUTION: Known only from the Northern Territory (map 41).