Anchophthalmops brevipleurum Koch, 1956

Kamiński, Marcin Jan, 2013, Taxonomy and distribution of the Afrotropical genus Anchophthalmops Koch, 1956 with a key to species (Coleoptera: Tenebrionidae: Pedinini), Zootaxa 3709 (6), pp. 501-523 : 505-521

Anchophthalmops brevipleurum Koch, 1956

(Figs 8, 18, 47)

Anchophthalmops brevipleurum Koch, 1956: 174 .—Kulzer 1963: 382; Iwan 2001a: 358, 2002a: 53, 2002b: 226, 2010: 32; Banaszkiewicz 2007: 395.

Studied material. Holotype, male (TMNH): “ Holotype No: 3219 / Anchophthalmops / brevipleurum / Koch”, “Pungwe Bay / East Africa / 12.1903 / P. Krantz”, “ Selinus / sp. nov. / ♂”. Other material: 2 males (MNHN): “Luabo / P.E.A. / I.1958 / P. Usher”; 3 males and 15 females (RGC): “ Mozambique. Dist. Sofala. / Inhamitanga S.E. Inhamitanga Safaris. / Campamento de Chironde (12). / 24.12/ 03.01.2006. / Leg. Robiche G. & Camiade D.”, “Collection / Robiche G.”; 2 males and 2 females (MNHN): Museum Paris / Zambeze / Forêt d'Inhacoro / J. Surcouf 1929”, “Décembre”; female (MNHN): “ Mozambique / Inhacoro / Près Chemba Inhacoro”, “Muséum Paris / 1936 / B. Thénot”; 2 females (MNHN): “Muséum Paris / Zambeze / Env. de Chemba / Bas Sangadze / P. Lesne 1929”; female (MNHN): “Muséum Paris / Zambeze / Nova Choupanga près Chemba / P. Lesne 1929”; male (MNHN): “Museum Paris / Bassin Inf. Du Zambeze / Wallee du Muza / 32 Long. E 16 Lat. S / ( DE 1000 A 1120 D`ALT) / G. Yasse 1905”; male (TMNH): “Luabo P.E.A. / X.1957 / B. Stuckenberg”; male (MNHN): “A. C. Rocha”, “Quelimane / 1557”; female (MNHN): “Beira, / Portuguese / E. Africa. / C. D. Rudd. / 1907–192.”; male (ZMHB): “Coll. Stuhlmann / Quilimane / 6.II.1889.”; male (MNHN): “Namacurra / X.16– 22 P.E.A. / C. B. Hardenberg”, “Muséum Paris Mozambique ”.

FIGURES 17–20. Body habitus. A. bartolozzii (17), A. brevipleurum (18), A. foveipennis (19), A. gridellii (20).

Redescription. Habitus as in Fig. 18. Body length = 10.0– 13.5 mm. Elytra wider and longer then pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.4–2.5).

Dorsal side of head dull, with conspicuous punctures (less then single diameter apart). Frontoclypeal suture distinct. Clypeal emargination shallow (clypeal emargination width / depth ratio ca. 8.0). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.5–1.6). Length of antennae equal or slightly greater then pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.0–1.2). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd ca. 3.8; ratio length of antenna / width of 3rd antennomere ca. 12.0).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.6); dull, with fine punctures (1–3 diameters apart). Anterior pronotal angles sharp and strongly protruding towards apex (Fig. 8). Lateral margins of pronotal disc rounded. Lateral sides of pronotal disc with longitudinal depressions. Pronotal hypomera dull, with fine punctures (5 diameters apart).

Elytra oblong (elytra length / width ratio = 1.2–1.3). Elytral striae with punctures (1–2 diameters apart). Intervals dull, with relatively small punctures (3–4 diameters apart). Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 4.5).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae = 5.0–5.4).

Abdominal process of both sexes with tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite = 1.9–2.1). Male 1st and 2nd abdominal ventrites with rounded tubercles. Bordering of 5th abdominal ventrite interrupted, with punctures 3–4 diameters apart.

Male legs. Protarsi widened; protibiae (protibiae length / width ratio ca. 3.6; profemora length / width ratio ca. 4.0) as in A. uhligi (Fig. 45). Mesotibiae, mesofemora simple. Metafemora setose on ventral side. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.)

Female legs simple (protibiae length / width ratio ca. 4.8; profemora length / width ratio ca. 4.4).

Parameres slightly widened at the base; narrowed towards apex; length equal to 0.2 of the rest of aedeagal tegmen. Clavae strait.

Paraproct longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix without sclerite. Spermatheca as in A. maximus ( Fig. 12 ).

Distribution. This species has been collected in the following ecoregions of East Africa ( Mozambique): East African mangroves, Southern Zanzibar-Inhambane coastal forest mosaic, Zambezian and Mopane woodlands, Zambezian coastal flooded savanna ( Fig. 47 ).

Anchophthalmops foveipennis (Fairmaire, 1887) (Figs 6, 9, 10, 13, 19, 31, 34, 42, 44, 48)

Selinus foveipennis Fairmaire, 1887: 286 .—Gebien 1910b: 277, 1938: 298; Koch 1956: 268, Iwan 2001a: 361.

Anchophthalmops foveipennis (Fairmaire, 1887) .—Iwan 2002a: 53, Iwan 2002b: 227; Banaszkiewicz 2007: 395.

Studied material. Three specimens with “ type ” labels are available. Fairmaire (1887) did not specify the number of individuals on which he described this species, thus a lectotype designation is needed to fix the taxonomic status of the species. Lectotype designated here, male (MNHN): “ Type ”, “Makdischu”, “ Platynotus / foveipenn- / Fairm / Makdischu”, “Museum Paris / 1906 / coll. L. Fairmaire” ( Fig. 13 ). Paralectotypes, female (MNHN): “Muséum Paris / 1906 / Coll. L. Fairmaire”, “ Somali ”; female (MNHN): “Muséum Paris / 1906 / Coll. L. Fairmaire”. Other material: 2 females (RGC): “ Somalie. Mogadisho. / 06.VII.1990. / Sous pierre.”, “Collection / Robiche G.”; male and female (MNHN): “ Somalia Mogadiscio / 7ºKm- 22/4-5/5 /1984 / R. Mourglia legit”; female (MNHN): “Nevinson Coll. / 1918-14”; male (MRAC): “Coll. Mus. Tervuren / Somalie: Afgoi / 1-VIII-1977 / leg. Olmi”; male and 2 females (MNHN): “Muséum Paris / 1890 / Coll. S. de Marseul”; male (MIIZ): “Mus. Zool. Polonicum / Warszawa / 12/45”, “ Somali ”, Selinus / foveipennis Frm. / H. Gebien det. 1939”; male (HNHM): “ Selinus / foveipennis Frm. / Dr Z. Kaszab det.”, “ Somalia IT. / Cesare Lomi / IX-XI-1935”; male (MNHN): “ Somali ”, “Muséum Paris”; “ Platynotus foveipennis / Somalis ”, “Ex-Musaeo / E. Allard / 1899”, “Museum Paris / ex. Coll. / R. Oberthur”; male (MNHN): “Museum Paris / Somali / Révoil 1885”, “ Platynotus / foveipennis ”.

Redescription. Habitus as in Fig. 19. Body length = 13.0–16.0 mm. Elytra wider and longer then pronotum (width ratio elytra / pronotum ca. 1.2; length ratio elytra / the middle of pronotum = 2.3–2.5).

Dorsal side of head shiny, with conspicuous punctures (less then single diameter apart). Frontoclypeal suture fine. Clypeal emargination deep (clypeal emargination width / depth ratio ca. 6.0). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere ca. 1.5). Length of antennae equal or slightly greater then pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.0–1.2). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd ca. 2.9; ratio length of antenna / width of 3rd antennomere ca. 16.0).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.6); shiny, with fine punctures (3 diameters apart). Lateral margins of pronotal disc parallel at the basal half. Lateral sides of pronotal disc with longitudinal depressions. Pronotal hypomera shiny, with fine punctures (5 diameters apart).

Elytra oblong (elytra length / width ratio = 1.2–1.3). Elytral striae with punctures (1–2 diameters apart). Intervals shiny, with relatively small punctures (3–4 diameters apart). Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 6.1).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 4.5).

In both sexes abdominal process without tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite = 1.9–2.1). Male 1st and 2nd abdominal ventrites with longitudinal tubercle. 5th abdominal ventrite with complete bordering (Fig. 31), with punctures 2–3 diameters apart.

Male legs. Protarsi widened; protibiae (protibiae length / width ratio ca. 6.6; profemora length / width ratio ca. 4.4) as in Fig. 44. Mesotibiae simple; mesofemora with denticle (Fig. 6). Metafemora setose on ventral side. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.)

Female legs simple (protibiae length / width ratio ca. 6.6; profemora length / width ratio ca. 4.4).

Parameres slightly widened at the base; narrowed towards apex; length equal to 0.2 of the rest of aedeagal tegmen ( Fig. 9 ). Clavae strait ( Fig. 10 ).

Paraproct longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix without sclerite. Spermatheca as in A. maximus ( Fig. 12 ).

Note. The original label of the lectotype suggests that this species was described within the genus Platynotus Fabricius, 1801 ( Fig. 13 ). However, Fairmaire in 1887 clearly included this species in the genus Selinus .

Distribution. This species has been collected in the following ecoregions of East Africa ( Somalia): Hobyo grasslands and shrublands, Somali Acacia - Commiphora bushlands and thickets ( Fig. 48 ).

Anchophthalmops gridellii (Ferrer, 1995) (Figs 1, 20, 38, 43, 48)

Selinus gridellii Ferrer, 1995: 34 .

Anchophthalmops gridellii (Ferrer, 1995) .—Iwan 2002b: 227; Banaszkiewicz 2007: 395.

Studied material. Holotype, male (NHRS): “ Holotype / Selinus , gridellii n. sp., det / Julio Ferrer 1992”, “ Somalia / Ola Uager / oltze Giuba 13–15 VIII 70 / Leg. S.B.S”, “La Specola / Firenze / 8180”. Paratypes, 3 females: same labels as holotype except: “8181”, 8181, “8182”. Other material: male and female (TMNH): “Kiunga / Mwini Isl. / July–Aug. 1951 / T. Huxley”; 4 males and female (MNHN): “Zanguebar / R. P. Le Roy / 1885.”, “Muséum Paris”; male and 3 females (MNHN): “Afr. O. A. / Kismayou”, “Museum Paris / Coll. Ch Allaud / 1925”; male (MNHN): “Afr. O. A. / entre Kismayou / et Afmadon”, “Museum Paris / Coll. Ch Allaud / 1925”; male (ZMHB): “Afr. or. / Lamu”; 14 males and 8 females (ZMHB): “O. affrica / Insel Lamri / Voelh Row J.”.

Morphological data. During the description (Ferrer 1995) this species was placed in Selinus Mulsant et Rey 1853 sensu Koch 1956 . Since that time, most of the species included in that genus were transferred to (Iwan 2002, Kamiński 2012): (1) Anchophthalmops ( A. menouxi , A. foveipennis ), (2) Angolositus Koch, 1953 ( A. elevatus (Gerstaecker, 1871) ), (3) Platykochius ( P. infernalis (Gerstaecker, 1854) , P. laekeyi (Koch, 1956) , P. platessa (Fairmaire, 1887)) and (4) Pseudoselinus Iwan, 2002 ( P. basilewskyi (Koch, 1956) , P. punctatostriatus (Gerstaecker, 1854) , P. raposoi (Koch, 1956)) . Thus, the original description of A. gridellii was adjusted to different taxonomic “surrounding”. Therefore, I propose to rewrite it in order to make it compatible with other data concerning the genus Anchophthalmops.

Habitus as in Fig. 20. Body length = 12.0–15.0 mm. Elytra wider and longer then pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.6–2.8).

Dorsal side of head dull, with conspicuous punctures (less then single diameter apart). Frontoclypeal suture visible. Clypeal emargination deep (clypeal emargination width / depth ratio ca. 4.0). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere ca. 1.0). Length of antennae greater then pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.2–1.4). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd ca. 3.0; ratio length of antenna / width of 3rd antennomere ca. 12.0).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.6); dull, with fine punctures (2–4 diameters apart). Anterior pronotal angles rounded and moderately protruding towards apex. Lateral margins of pronotal disc rounded. Lateral sides of pronotal disc with longitudinal depressions. Pronotal hypomera dull, with fine punctures (5 diameters apart).

Elytra oblong (elytra length / width ratio = 1.3–1.5). Elytral striae with punctures (1–2 diameters apart). Intervals dull, with relatively small punctures (3–4 diameters apart). Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 5.8).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae = 4.5–5.0).

In both sexesabdominal process without tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite ca. 2.0). Male 1st and 2nd abdominal ventrites with rounded tubercles (Fig. 38). 5th abdominal ventrite with complete bordering, with punctures 1–2 diameters apart.

Male legs. Protarsi widened; protibiae as in Fig. 43 (protibiae length / width ratio = 5.5–5.6; profemora length / width ratio = 3.5–4.0). Mesotibiae simple; mesofemora with denticle. Metafemora setose on ventral side. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.)

Female legs simple (protibiae length / width ratio = 5.5–5.6; profemora length / width ratio = 3.5–4.0).

Parameres slightly widened at the base; narrowed towards apex; length equal to 0.2 of the rest of aedeagal tegmen. Clavae strait.

Paraproct longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix without sclerite. Spermatheca as in A. maximus ( Fig. 12 ).

Distribution. This species has been collected in the following ecoregions of East Africa ( Kenya, Somalia, Zanzibar): East African mangroves, Northern Zanzibar-Inhambane coastal forest mosaic, Somali Acacia - Commiphora bushlands and thickets ( Fig. 47 ).

Anchophthalmops maximus Koch, 1956 (Figs 7, 11, 12, 21, 36, 37, 46)

Anchophthalmops maximus Koch, 1956: 178 .—Kulzer 1963: 382; Iwan 2001a: 358, 2002a: 53, 2002b: 227; Banaszkiewicz 2007: 395.

Studied material. Holotype, male (TMNH): “Musée du Congo / B.E.A.: Merifano / IX. 1932 / C.G. Macarthur”, “Holo / type / maximus / Koch”. Paratypes: female (MGFT): “ Paratypus ”, “Musée du Congo / B.E.A.: Nibulola / 6-V-1913 / Dr. Bayer”; female (TMNH): “ Paratypus ”, “Musée du Congo / B.E.A.: Nibulola / 6-V-1913 / Dr. Bayer”; female (TMNH) “ Paratypus ”, “Musée du Congo / B.E.A.: Zuwani / 28-VI-1913 / Dr. Bayer”, “ Anchophthalmops maximus Koch / C. Koch det., Koch 1956”. Other material: female (SMNS): “Deutach– Ostafrica / Kilimandacharo / 1931–1934 / W. Othmöller leg.”; 2 females (MNHN): “Museum Paris / Afrique Orient. Angl. / Sambourou / Wanyika / Ch. Alluaud 1904”, “P 0 0 0 996”; 5 males and 7 females (MNHN): “Museum Paris / Afrique Orient. Angl. / Pori de Serenheti / De Boura a Taveta / Ch. Alluaud 1904”, “P 0 0 0 999”; male (MZUF): “ Somalia: Showli Berdi / esterno: sprofondamento / ficus ramo Mugdile / 27.XI.1985 S.B.S!”, “La Specola / Fienze / 8158”; 2 males and 2 females (RGC): “ Kenya. Hola. E. prov. / 13.V.2001 / leg. K. Werner.”, “Collection Robiche Gérard”.

Redescription. Habitus as in Fig. 21 . Body length = 14.0–22.0 mm. Elytra wider and longer then pronotum (width ratio elytra / pronotum = 1.1–1.3; length ratio elytra / the middle of pronotum = 2.3–2.4).

Dorsal side of head dull, with conspicuous punctures (0.5–1 diameters apart). Frontoclypeal suture distinct. Clypeal emargination shallow (clypeal emargination width / depth ratio = 5.4–5.6). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.2–1.3). Length of antennae equal or slightly greater then pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.0–1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 4.6–5.0; ratio length of antenna / width of 3rd antennomere ca. 13.5).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.6); dull, with fine punctures (4–5 diameters apart). Anterior pronotal angles rounded and slightly protruding towards apex. Lateral margins of pronotal disc rounded. Lateral sides of pronotal disc with longitudinal depressions. Pronotal hypomera shiny, with fine punctures (5 diameters apart).

Elytra oblong (elytra length / width ratio = 1.2–1.3). Elytral striae with conspicuous punctures (0.5–1 diameters apart). Intervals dull, with relatively small punctures (4–5 diameters apart). Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 5.0).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae = 4.1–4.3).

In both sexes abdominal process with tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite = 2.1–2.2). Male 1st and 2nd abdominal ventrites with tubercles. 5th abdominal ventrite with complete bordering, with punctures 2–3 diameters apart.

Male legs. Protarsi widened; protibiae as in Fig. 36 (protibiae length / width ratio = 3.5–3.6; profemora length / width ratio = 4.6–4.8). Mesotibiae with denticles situated at 2/3 of their length (Fig. 37). Meso- and metafemora simple. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.)

Female legs simple (protibiae length / width ratio = 3.5–3.6; profemora length / width ratio = 4.6–4.8).

Parameres slightly widened at the base; narrowed towards apex; length equal to 0.2 of the rest of aedeagal tegmen. Clavae strait.

Paraproct longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix without sclerite. Spermatheca as in Fig. 12 .

Distribution. This species has been collected in the following ecoregions of East Africa ( Kenya, Somalia, Tanzania): East African montane forests, Northern Acacia - Commiphora bushlands and thickets, Northern Zanzibar-Inhambane coastal forest mosaic, Serengeti volcanic grasslands, Somali Acacia - Commiphora bushlands and thickets ( Fig. 46 ).

Anchophthalmops menouxi (Mulsant et Rey, 1853) (Figs 5, 22, 27, 29, 49, 50)

Selinus menouxi Mulsant et Rey 1853: 322 .—Gemminger and Harold 1870: 1915; Gebien 1910b: 278, 1938: 298; Koch 1956: 260; Iwan 1995: 52.

Anchophthalmops menouxi (Mulsant et Rey 1853) .—Iwan 2002a: 53, 2002b: 227; 2010: 33; Banaszkiewicz 2007: 395.

Opatrinus sayi Horn, 1870: 349 .—Crotch 1873: 106; Henshaw 1885: 119; Gebien 1910b: 277, 1938: 296; Papp 1961: 119; Iwan 1995: 52 (syn.).

Opatrinus trivialis Gerstaecker, 1871: 60 .—Gerstaecker 1873: 177; Koch 1956: 260 (syn.).

Selinus trivialis (Gerstaecker, 1871) .—Fairmaire 1887: 284; Gebien 1910b: 278, 1938: 298.

Studied material. Lectotype [designated by Iwan in 2010], male (MNHN): “Afrique, coll. Chevrolat”. Paralectotypes, 2 males (MNHN); “ Selinus Menouxi, Madag., Muls. ”. Other material. 3 males and 2 females (MNHN): “Bagamoyo / H. Schaedle”, “Museum Paris”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and female (MNHN): “Ost-Africa / Dar-es Salaam / ex coll. Fruhstorfer”, “Muséum Paris / 1906 / Coll. L. Fairmaire”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; female (ZMHB): “ Witu / Kinini / F. Thomas S. / G. Denhardt G.”; female (ZMHB): “ Witu / Pokomonie / G. Denhardt G.”; female (ZMHB): “D. Ost- Africa / Korogwe / Mitte V. 93. / O. Neumann S.”; male 3 females (ZMHB): “D. Ost-Africa / Tan?a, Um?e?end / III-IV. 93. / O. Neumann S.”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; female (ZMHB): “D. Ost-Africa / Tan?a / Ende IV. 93. / O. Neumann S.”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; female (ZMHB): “D. Ost-Africa / Tan?a / Ende IV. 93. / O. Neumann S.”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male (ZMHB): “Brit. Ost-Africa / Jnsel Pamba / Chake-Chake / 14.IV.1903 / Voeltzkow S.”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 5 males and 4 females (MNHN): “MUSEUM PARIS / OUSAMBARA / ENV. DE TANGA / GIERRA 98-97”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 3 males and 7 females (MRAC): “Coll. Mus. Tervuren / Tanganyika: Manow / ex. Coll. Breuning”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and 2 females (MNHN): “MUSEUM PARIS / AFRIQUE ORIENTALE / TANGA / GIERRA 1895”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 2 males and 4 females (ISNB): “ Zanzibar / Raffray”, “R. I. Sc. N. B. / I. G. 23.107”, “Coll. D. / Ach. 1.90”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 4 females (ZMHB): “Pemba, Jnsel / [illegible] / IV.03 / Voelhkow J.”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and 2 females (ZMHB): “N. Victoria-Nyansa / Ussoga u. Uganda / V.-VIII. 1894 / O. Neumann S.”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and 2 females (MRAC): “Silenus / trivialis / Gst.”, “Dtsch. O. / Africa”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 2 males (MNHN): “MUSEUM PARIS / SOMALI / RÉVOIL1885”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 2 females (MNHN): “MUSEUM PARIS / KONDOA / BLOYET”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and female (ZMHB): “e. coll. Dr. C. Schroder / Magamba —Bge. / bei Masinde 6.I.05 / 1600-2000 m D.O. Afr”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and 6 females (MNHN): “MUSEUM PARIS / CONGO / DYBOWSKI 128-96”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 9 males and 14 females (ZMAS): “Tanga, Deutsch- / Ostafrica, Küste. / Gierra.”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 4males and 4 females (MNHN): “ Zanzibar / R.P. Guillemé”, “MUSEUM PARIS”; 2 males and 3 females (ISNB): “ Zanzibar / Raffray”, “R. I. Sc. N. B. / I. G. 23.107”, “Coll. D. / Ach. 1.90”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and female (MNHN): “Zangebar. / R. P. Leroy.”, “MUSÉUM PARIS / 1952 / coll R. Oberthur”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 2 males and female (MNHN): “Coll. Stuhlmenn / Bagamoyo / Febr. 1890”, “ Selinus trivialis / Gst.”; 3 males (MIIZ): “ Zanzibar, / C. Cooke.”, “ Selinus trivialis / H. Gebien 1939”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 2 females (MNHN): “Côte d`Afrique or angl. / Shimoni / Alluaud et Jeannel / Nov. 1911 St. 9”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and female (MNHN): “Côte d`Afrique or angl. / Riviére Rimisi / Alluaud et Jeannel / Nov. 1911 St. 5”, “MUSÉUM PARIS”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and 2 females (MNHN): “Côte d`Afrique or angl. / Ile de Mombasa / Alluaud et Jeannel / Nov. 1911 St. 5”, “MUSÉUM PARIS”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; 3 males (HNHM): “ TANZANIA, Tanga reg. / Chemka, lateral valley / of Hunga river”, “swept & beaten, No. 88, / 10. II. 1987 / S. Mahunka & A. Zicsi”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and 2 females (Lillig collection): “ Selinus ? menouxi / Mulsant et Rey / det. M. Lillig, 2005”, “ Tanzania / Ulunguru Mts. / III.1991 / leg. E. Rautenstrauch”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; female (BMNH): “Brit. E. Africa. / Magogoni swamp, / near Witu / 29 Feb. 1912 / S.A. Neave.”; male (BMNH): “Brit. E. Africa. / Kipini, on coast, / at mouth of Tana. / near Witu / 3 March 1912 / S.A. Neave.”; female (BMNH): “Brit. E. Africa. / Uchweni forest, / near Witu / 25–27 Feb. 1912 / S.A. Neave.”; male (BMNH): “Brit. E. Africa. / Voi. / 8–10 Feb. 1912 / S.A. Neave.”; female (BMNH): “Brit. E. Africa. / Mombasa. / 12–13 Feb. 1912 / S.A. Neave.”; male (ZMHB): “D.O. AFRICA /? Muansa / Dr. M. Schmidt”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male (BMNH): “V.G.L. Van Someren / Collection. / Brit. Mus. 1959–468.”, “COM. INST. ENT. / COLL NO. 10829”, “van Someren / Tiwi / March 1941 ”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; female (MNHN): “Turiani T.T. / N. Morogoro / Apr 1 51 / E. Pinhey”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; female (MIIZ): “ 30.IX.1985. leg. / S. & L. Mahunka”, Diani Persian / Mosque,”, “ KENYA No. 85 / Ukunda”, “pitfall / trap”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male (MIIZ): “ 18.IX.1985. leg. / S. & L. Mahunka”, Diani Persian / Mosque,”, “ KENYA No. 26 / Kwale”, “pitfall / trap”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male (MIIZ): “ KENYA, Ukunda, / Diani Beach, singled / No. 163, 2.VII.1992 / S. & L. Mahunka”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male and female (HNHM): “D.O. Africa / Nguru”, “ trivialis Gerst. / det. dr. Kaszab”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male (MIIZ): “Tanganyka / Dr. Szunyoghy”, “Tanga / 25–26.III.1960 ”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male (MNHN): “Zanbuebar / Mhonda-Ouzigoua / A. Hacquard Mis. ap. / 1879 1st Trim. 1880”; female (SMNS): “ Tansania Tanga / 21.5.81 / leg. M. Hemdrich”; male (ZMHB): “ Selinus / trivialis / Gest.”; male and female (MIIZ): “ex. Coll. R. B.ennigsen / Inst. Zool. P.A.N. / Warszawa / 52 / 58”, “Coll. v. Bennigsen”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; female (MRAC): “Coast Prov.: Shimba / Hills, Kwale alt. / 400m / 25-VII-1977 / G. Coulon”, “Coll. Mus. Tervuren / Kenya: VII/ IX-1977 / G. Coulon”; male (MNHN): “Côte d`Afrique or angl. / Cheteni. / Alluaud et Jeannel / Nov. 1911 St. 9”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”; male (MNHN): “ MADAGASCAR / COLL. SCHRAMM”, “MUSEUM PARIS / COLL. J. CHATANAY 1914”, “ Anchophthalmops / menouxi (Muls. et Rey) / det. D. Iwan”.

Redescription. Habitus as in Fig. 22 . Body length = 10.0–14.0 mm. Elytra wider and longer then pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.5–2.7).

Dorsal side of head dull, with conspicuous punctures (0.5–1 diameters apart). Frontoclypeal suture visible. Clypeal emargination shallow (clypeal emargination width / depth ratio ca. 5.6). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.1–1.2). Length of antennae equal or slightly greater then pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.0–1.1). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.9–3.4; ratio length of antenna / width of 3rd antennomere ca. 11.5).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.7); dull, with fine punctures (3–4 diameters apart). Anterior pronotal angles rounded and slightly protruding towards apex (Fig. 7). Lateral sides of pronotal disc with longitudinal depressions. Pronotal hypomera dull, without punctures.

Elytra oblong (elytra length / width ratio = 1.3–1.4). Elytral striae with fine irregularly distributed punctures. Intervals dull, without punctures. Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 5.0).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 4.5).

In both sexes abdominal process without tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite = 2.1–2.2). Male 1st and 2nd abdominal ventrites with rounded tubercles. 5th abdominal ventrite bordered; with fine punctures 1–2 diameters apart.

Male legs. Protarsi widened; protibiae (protibiae length / width ratio = 4.8–5.2; profemora length / width ratio = 3.6–4.0) as in A. uhligi , see Fig. 45. Mesotibiae simple; mesofemora with denticle (as in A. foveipennis , see Fig. 6). Metafemora setose on ventral side. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.)

Female legs simple (protibiae length / width ratio = 4.8–5.2; profemora length / width ratio = 3.6–4.0).

Parameres slightly widened at the base; narrowed towards apex; length equal to the 0.2 of the rest of aedeagal tegmen. Clavae strait.

Paraproct longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix without sclerite. Spermatheca as in A. maximus ( Fig. 12 ).

Distribution. This species has been collected in the following ecoregions of East Africa ( Kenya, Tanzania, Uganda): Cental Zambezian Miombo woodlands, East African mangroves, Northern Zanzibar-Inhambane coastal forest mosaic, Northern Acacia - Commiphora bushlands and thickets, Southern Acacia - Commiphora bushlands and thickets, Southern Rift montane forest-grassland mosaic, Victoria Basin forest-savanna mosaic ( Fig. 49 ). Modeled species range is presented on Fig. 50 .

Remarks. Labels of some of the studied specimens as well as available published data (Gebien 1938, Iwan 2010) implied that Madagascar is a part of the distributional range of A. menouxi . However, none of these sources specify the exact locations within the island.

The issue of the hypothetical occurrence of A. menouxi on Madagascar was briefly discussed by Iwan in 2010. In his opinion, the eastern part of Africa is the main part of the distributional area of the discussed species —and the whole genus Anchophthalmops —therefore finding introduced specimens on Madagascar is possible. Unfortunately, the studies of the diverse entomological material —concerning the tribe Pedinini —acquired from California Academy of Sciences expeditions to Madagascar between 2000 and 2002, did not confirmed Iwan`s suggestion (Iwan et al. 2011, 2012; Iwan and Kamiński 2012; Kamiński and Raś 2012).

Because of a relatively high diversity of entomological material, representing A. menouxi , it was possible to model a species range and estimate the probability of the occurrence of this species on Madagascar. The results obtain in the MaxEnt software confirms Iwan`s suggestion that the eastern part of central Africa is the main distributional area of A. menouxi . However, according to the predictive distribution model it is unlikely for A. menouxi to occur on Madagascar ( Fig. 50 ).

In my opinion, the lack of the unambiguous “label evidence” as well as the estimated MaxEnt model proves that the registered occurrence of A. menouxi on the Madagascar was caused by mislabeling.

Within the whole Ectateus generic group only species of Lechius Iwan 1995 occur on Madagascar and all of five species of this taxon are endemic (Iwan 2001b, Iwan and Kamiński 2012). Therefore, the Mozambique Channel seems to be efficient dispersion barrier of the representatives of Ectateus generic group.

Anchophthalmops neumanni sp. nov. ( Figs 15 , 23 , 28, 30 , 33, 41, 47)

Type data. Holotype, male (ZMHB): “Tabora” ( Fig. 15 ).

Locus typicus. Tabora ( Tanzania).

Etymology. I dedicate this species to my dear friend Adam Karol Neumann (Warsaw, Poland).

Diagnosis. This species is similar to A. menouxi in the structure of 5th abdominal ventrite (completely bordered), elytral striae impressed on the whole length (Fig. 33), male abdominal process (without tubercle) and male protibia (Fig. 45). The two species can be distinguished by the different body size ( A. neumanni : 15.0 mm; A. menouxi : 10.0–14.0 mm), structure of pronotum (strongly transverse in A. neumanni — Fig. 30 ), elytral epipleuron (9th elytral striae invisible from the ventral side in A. neumanni — Fig. 28 ) and elytral striae (with coarse regularly distributed punctures in A. neumanni ).

Description. Habitus as in Fig. 23 . Body length ca. 15.0 mm. Elytra wider and longer then pronotum (width ratio elytra / pronotum ca. 1.1; length ratio elytra / the middle of pronotum ca. 2.8).

Dorsal side of head dull, with conspicuous punctures (0.5–1 diameters apart). Frontoclypeal suture visible. Clypeal emargination shallow (clypeal emargination width / depth ratio ca. 7.5). Mentum with median part wide. Submentum with short base. 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd ca. 2.6).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.5); dull, with visible punctures (2–4 diameters apart). Anterior pronotal angles rounded and moderately protruding towards apex ( Fig. 30 ). Basal half of lateral sides of pronotal disc straight. Pronotal hypomera dull, without punctures.

Elytra oblong (elytra length / width ratio ca. 1.2). Elytral striae with conspicuous punctures (0.5–1 diameter apart). Intervals dull, with fine punctures (4–5 diameters apart). Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Epipleura without punctation. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 4.7).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 4.1).

Male abdominal process without tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite ca. 2.0). Tubercles on 1st and 2nd abdominal ventrites longitudinal. 5th abdominal ventrite bordered; with punctures 1 diameters apart.

Male legs. Protarsi widened; protibiae (protibiae length / width ratio ca. 4.7; profemora length / width ratio ca. 4.5) as in A. uhligi , see Fig. 45. Mesotibiae simple; mesofemora with a denticle (as in A. foveipennis , see Fig. 6). Metafemora setose on ventral side. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.).

Parameres slightly widened at the base; narrowing towards apex; length equal to the 0.2 of the rest of aedeagal tegmen. Clavae strait.

Notes. This species is known only from one specimen (male).

Distribution. This species has been collected in the following ecoregion of East Africa ( Tanzania): Hobyo grasslands and shrublands ( Fig. 47 ).

Anchophthalmops ventralis (Gebien, 1910) ( Figs 25 , 40, 46)

Selinus ventralis Gebien, 1910a: 375 .—Gebien 1910b: 278, 1938b: 298.

Anchophthalmops ventralis (Gebien, 1910a) .—Koch 1956: 176; Kulzer 1963: 382; Iwan 2001a: 358, 2002a: 53, 2002b: 228; Banaszkiewicz 2007: 395.

Studied material. Two specimens with “ type ” labels are available. Gebien (1910a) did not specify the number of individuals on which he described this species, thus a lectotype designation is needed to fix the taxonomic status of the species. Lectotype designated here, male (MGFT): “Dar es Sal.”, “ Type: / No. 89”, “ Selinus / ventralis / X Feb”, “Anchophthalmops / ventralis (Gebien) / det. D. Iwan”, “ Lectotypus ” ( Fig. 14 ). Paralectotype, female (MGFT): “Usambara / Mombo / (Sjoestedt)”, “ Type! / No. 89”, “ Selinus ventralis / x Geb”, “Anchophthalmops / ventalis (Gebien) / det. D. Iwan”, “ Paralectotypus ”. Other material: male and female (ZMHB): “D. Ost-Africa / Usambara, Kwai / P. Weise S.”, male (MIIZ): “ Selinus / ventralis Geb. / det. dr. Kaszab”.

Redescription. Habitus as in Fig. 24 . Body length = 14.0–17.0 mm. Elytra wider and longer then pronotum (width ratio elytra / pronotum = 1.1–1.2; length ratio elytra / the middle of pronotum = 2.3–2.5).

Dorsal side of head dull, with conspicuous punctures (0.5–1 diameters apart). Frontoclypeal suture distinct. Clypeal emargination shallow (clypeal emargination width / depth ratio ca. 6.4). Mentum with median part wide. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.2–1.3). Length of antennae equal or slightly greater then pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.0–1.1). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8–3.2; ratio length of antenna / width of 3rd antennomere ca. 14.5).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.7); dull, with fine punctures (4–5 diameters apart). Anterior pronotal angles rounded and moderately protruding towards apex. Lateral margins of pronotal disc parallel at the basal half. Lateral sides of pronotal disc with longitudinal depressions. Pronotal hypomera dull, with fine punctures (5 diameters apart).

Elytra oblong (elytra length / width ratio = 1.2–1.3). Elytral striae with conspicuous punctures (2–3 diameters apart). Intervals dull, with relatively small punctures (3–4 diameters apart). Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 5.0).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae = 5.7–5.9).

In both sexes abdominal process without tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite = 2.1–2.2). Male 1st abdominal ventrite with transverse tubercle (Fig. 39); 2nd abdominal ventrite with rounded tubercle. Bordering of 5th abdominal ventrite interrupted (Fig. 32), with punctures 2–3 diameters apart.

Male legs. Protarsi widened; protibiae (protibiae length / width ratio = 4.0–5.8; profemora length / width ratio = 3.2–3.5) as in A. maximus (Fig. 36). Mesotibiae with denticles situated at 2/3 of their length. Meso- and metafemora simpe. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.)

Female legs simple (protibiae length / width ratio = 4.0–5.8; profemora length / width ratio = 3.2–3.5).

Parameres slightly widened at the base; narrowed towards apex; length equal to the 0.2 of the rest of aedeagal tegmen. Clavae strait.

Paraproct longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix without sclerite. Spermatheca as in A. maximus ( Fig. 12 ).

Distribution. This species has been collected in the following ecoregion of East Africa ( Tanzania): Northern Zanzibar-Inhambane coastal forest mosaic ( Fig. 46 ).

Anchophthalmops uhligi Banaszkiewicz, 2007 (Figs 3, 4, 14, 24, 32, 39, 45, 47)

Anchophthalmops uhligi Banaszkiewicz, 2007: 400 .

Studied material. Holotype, male (HNHM): “ Holotype; Anchophthalmops uhligi sp. nov. det. M. Banaszkiewicz 2007 / Selinus planipleurus aff. infernalis det. dr. Kaszab / O. Africa, Ikulo”. Paratypes, male (HNHM): “ Selinus planipennis det. Dr. Kaszab, Sammlung Hans Ertl”; male (ZMHB): “Deutsch-Ostafrica H. Dabbert”; female (ISNB): “Africa or. Micindani, F. Schneider / ex. Coll. Cl. Segers”. Other material: male (ZMHB): “D.O.-Africa / Lindi / O. Werner S.”; female (MNHN): “Afr. or. all. / Lukuledi”, “Muséum Paris”.

Morphological data. Because the original description (Banaszkiewicz 2007) of this species is relatively recent and consistent with the description style adopted in this study the morphology of this species was not redescribed.

Distribution. This species has been collected in the following ecoregions of East Africa ( Tanzania): Central Zambezian Miombo woodlands, Eastern Miombo woodlands, Southern Zanzibar-Inhambane coastal forest mosaic ( Fig. 47 ).

Anchophthalmops zablockii sp. nov. ( Figs 16 , 26 , 35, 46)

Type data. Holotype, male (HNHM): “ Somalia V.1980 / Afgoi (Mogadiscio) / leg. Olmi”, “ Selinus (olim) / det. Kaszab sp.”, “ Selinus / n. sp.” ( Fig. 16 ).

Locus typicus. Mogadishu ( Somalia).

Etymology. I dedicate this species to my dear friend Wojciech Zabłocki (Warsaw, Poland).

Diagnosis. This species is similar to A. maximus in the body size, structure of protibia, mesotibia and 5th abdominal ventrite (completely bordered). The two species can be distinguished by the different occurrence of tubercles on the abdominal ventrites (lack of tubercles on the apex of 1st and 2nd segments in A. zablockii sp. nov.) (Fig. 35). The two species differ also in the structure of elytral surface (shiny in A. zablockii sp. nov. — Fig. 26 , dull in A. maximus — Fig 21 ).

Description. Habitus as in Fig. 26 . Body length ca. 20.0. Elytra wider and longer then pronotum (width ratio elytra / pronotum ca. 1.2; length ratio elytra / the middle of pronotum ca. 2.0).

FIGURES 31–37. 5th abdominal ventrite (31, 32), surface of elytral disc (33, 34), 1st and 2nd abdominal ventrite (35), male pro- (36) and mesotibia (37). A. foveipennis (31, 34), A. ventralis (32), A. neumanni sp. nov. (33), A. zablockii sp. nov. (35), A. maximus (36, 37).

FIGURES 38–45. 1st and 2nd abdominal ventrites (38, 39), elytral humerus (40), mesotibia (41), elytral apex (42), protibia (43, 44, 45). A. gridellii (38, 43), A. ventralis (39), A. uhligi (40, 45), A. neumanni (41), A. foveipennis (42, 44).

Dorsal side of head dull, with conspicuous punctures (0.5–1 diameters apart). Frontoclypeal suture distinct. Clypeal emargination shallow (clypeal emargination width / depth ratio ca. 5.0). Mentum with median part wide. Submentum with short base. 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd ca. 3.0).

Pronotal disc transverse (middle of pronotum length / width ratio ca. 0.6); dull, with fine punctures (4–5 diameters apart). Anterior pronotal angles rounded and slightly protruding towards apex. Lateral sides of pronotal disc rounded. Lateral sides of pronotal disc without longitudinal depressions. Pronotal hypomera shiny, with fine punctures (5 diameters apart).

Elytra oblong (elytra length / width ratio ca. 1.2). Elytral striae with conspicuous punctures (1 diameter apart). Intervals shiny, with fine punctures (4–5 diameters apart). Elytral base sinusoidal. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; relatively narrow (width ratio anterior margin of elytra / scutellum at base ca. 3.7).

Intercoxal process of prosternum straight with complete bordering. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae = 5.7–5.9).

Male abdominal process with tubercle; relatively narrow (process of I abdominal ventrite / process of metaventrite ca. 2.0). Male 1st and 2nd abdominal ventrites without tubercles. 5th abdominal ventrite bordered, with punctures 1–3 diameters apart.

Male legs. Protarsi widened; protibiae (protibiae length / width ratio ca. 4.7; profemora length / width ratio ca. 4.7) as in A. maximus (Fig. 36). Mesotibiae with denticles situated at 2/3 of their length. Meso- and metafemora simpe. 1st metatarsomere elongated (length ratio metatarsomere 1st / 2nd ca. 2.0.).

Parameres slightly widened at the base; narrowed towards apex; length equal to 0.2 of the rest of aedeagal tegmen. Clavae strait.

Notes. This species is known only from one, partly damaged, specimen (male). The structure of antennal flagellum, distal segment of the maxillary palpi, pro-, mesotarsi and hind legs is unknown. Distribution. This species has been collected in the following ecoregion of East Africa ( Somalia): Hobyo grasslands and shrublands ( Fig. 46 ).

Key to the species of Anchophthalmops:

1. 1st abdominal ventrite completely bordered (Fig. 1)........................................................... 2

- Bordering of the 1st abdominal ventrite interrupted (Fig. 2)..................................................... 7

2. Elytral striae impressed on whole length (Fig. 3)............................................................. 3

- Elytral striae impressed only near the punctures (Fig. 4)...................................................... 5

3. In both sexes abdominal process with rounded tubercle. Male protibiae as in Fig. 36; mesotibiae with denticles situated at 2/3 of its length (Fig. 37).......................................................................... A. maximus

- Abdominal process without tubercle. Male protibia as in A. uhligi , see Fig. 45; mesotibia simple (Fig. 41)............... 4

4. Body size: 10.0–14.0 mm. 9th elytral striae invisible from the ventral side ( Fig. 27 ). Elytral striae with fine irregularly distrib- uted punctures. Pronotum slightly transverse (middle of pronotum length / width ratio ca. 0.7) ( Fig. 29 )........ A. menouxi

- Body size: 15.0 mm. Part of 9th elytral row visible from the ventral side ( Fig. 28 ). Elytral striae with coarse regularly distrib- uted punctures (0.5–1 diameters apart). Pronotum strongly transverse (middle of pronotum length / width ratio ca. 0.5) ( Fig. 30 )................................................................................ A. neumanni sp. nov.

5. Body size: 20.0 mm. Abdominal process with rounded tubercle (Fig. 35). Male 1st and 2nd abdominal ventrites without tuber- cles; protibiae as in A. maximus (see Fig. 36)................................................ A. zablockii sp. nov.

- Body size: 13.0–16.0 mm. Abdominal process without tubercle. Male 1st and 2nd abdominal ventrites with tubercles; protibiae as in Figs. 43, 44..................................................................................... 6

6. Elytral apices contacting. Lateral sides of pronotum rounded. Pronotal disc dull............................ A. gridellii

- Elytra diverge just before apex (Fig. 42). Lateral sides of pronotum parallel at the basal half. Pronotal disc shiny..................................................................................................... A. foveipennis

7. Anterior pronotal angles sharp and strongly protruding towards apex (eg Fig. 8). Male protibiae as in Fig. 45; mesotibiae sim- ple. Male abdominal process with tubercle; 1st abdominal ventrite with rounded tubercle............................ 8

- Anterior pronotal angles rounded and moderately protruding towards apex. Male protibiae as in A. maximus (see Fig. 36); mesotibiae with denticles situated at 2/3 of its length. Abdominal process without tubercle; 1st abdominal ventrite with trans- verse tubercle (eg Fig. 39).............................................................................. 9

8. Body size 15.0–20.0 mm. Elytral humeri protruding laterad (Fig. 40). Female abdominal process without tubercle... A. uhligi

- Body size 10.0– 13.5 mm. Elytral humeri not protruding laterad. In both sexes abdominal process with rounded tubercle............................................................................................ A. brevipleurum

9. Body size 10.5–11.5 mm. Lateral pronotal sides narrowing toward apex................................ A. bartolozzii

- Body size 14.0–17.0 mm. Lateral pronotal sides parallel at basal half.................................... A. ventralis

Figures

FIGURE 50. Modeled range of A. menouxi generated in MaxEnt. Different hues of green indicate the probability of occurrence in percent. The model was calculated with exclusion of locations situated closer than 20 km from each other. White dots indicate the documented species distribution. Training AUC = 0.946.

FIGURES 9 – 16. Aedeagal tegmen dorsal and ventral view (10, 11), ovipositor with part of bursa copulatrix (11), spermatheca (12), labels of primary types (13 – 16). A. foveipennis (9, 10, 13), A. maximus (11, 12), A. ventralis (14), A. neumanni sp. nov. (15), A. zablockii sp. nov. (16).

FIGURES 46 – 49. Distribution of the species of Anchophthalmops. The division of Afrotropical Realm into ecoregions was adopted after Olson et al. 2001. Different colors were used to distinguish the adjacent ecoregions.

FIGURES 21 – 24. Body habitus. A. maximus (21), A. menouxi (22), A. neumanni sp. nov. (23), A. ventralis (24).

FIGURES 25 – 30. Body habitus (25, 26), epipleuron (27, 28), pronotal disc (29, 30). A. uhligi (25), A. zablockii sp. nov. (26), A. menouxi (27, 29), A. neumanni sp. nov. (28, 30).