Sahastata wunderlichi,

Magalhaes, Ivan L. F., Stockmann, Mark, Marusik, Yuri M. & Zonstein, Sergei L., 2020, On Sahastata (Araneae: Filistatidae): complementary description of the generotype and two new species from Oman and Morocco, Zootaxa 4899 (1), pp. 215-246: 235-242

publication ID

publication LSID


persistent identifier

treatment provided by


scientific name

Sahastata wunderlichi

sp. nov.

Sahastata wunderlichi  sp. nov.

Figures 1View FIGURE 1 E–F, 18–23, 24C– D

Sahastata nigra Benoit, 1968: 97  , figs 1–5 (♀ misidentified in part, figures refer to a species near S. wunderlichi  sp. nov.; material examined may refer in part to S. wunderlichi  sp. nov., in part to S. infuscata  , and in part to undescribed species); Marusik et al. 2014: 9, figs 30–33, 41 (♀, fig. 41 copy of fig. 4 in Benoit 1968 — Sahastata cf. wunderlichi  ).

Remarks. Specimens examined and listed by Benoit (1968) from Morocco that were identified as S. nigra  seem to refer to S. wunderlichi  sp. nov..

Etymology. The name honours Jörg Wunderlich, the first to study the holotype of S. bosmansi  , in recognition of his life-long contributions to the systematics of extant and fossil spiders.

Diagnosis. In possessing a sideward-directed embolus, males of S. wunderlichi  sp. nov. are similar to those of S. bosmansi  ; they differ by the ventral position of the embolic keel and the gentle curvature of the embolus ( Fig. 20View FIGURE 20) (retrolateral keel and sharp curve in S. bosmansi  ; Fig. 17View FIGURE 17). The spermathecae resemble those of S. amethystina  , but in S. wunderlichi  sp. nov. there is no membranous portion of the spermathecae, and the calamistrum has more setae in each row ( Figs 22BView FIGURE 22, 23View FIGURE 23).

Description. Male (Paratype, MACN-Ar 39465). Coloration pale yellowish cream, except where noted. Carapace with dark pigment around eyes and faint light brown median V-shaped pattern. Carapace with thick black setae interspersed with regular setae. Sternum with white markings posteriorly. Abdomen light yellowish brown.

Total length 4.94. Carapace 2.44 long, 1.96 wide. Clypeus 0.3 long. Eye sizes and interdistances: AME 0.13, PME 0.14, ALE 0.198, PLE 0.18, AME-AME 0.06, PME-PME 0.156. Sternum length 1.19, width 1.06. Palp: femur length 3.64, height 0.37, tibia length 3.23, height 0.33. Leg I: fe 5.12, pa 1.12, ti 5.08, mt 5.9, ta 2.28. II: fe 4.1, pa 0.95, ti 3.77, mt 4.37, ta 1.81. III: fe 3.8, pa 0.75, ti 3.27, mt 4.3, ta 1.55. IV: fe 4.99, pa 1.07, ti 4.79, mt 5.85, ta 2.46. Abdomen: length 2.68, width 1.16.

Palp with long and numerous macrosetae in several rows along femur ventral and dorsal sides. Leg macrosetae: fe I dp8, v31, with large patch with ~50 short retrolateral macrosetae; ti I r0-1-1, v2-2-2-2; mt I p0-1-0, v2-2-2-2- 2-2-2; ta I v4; fe II v23, with a large patch with ~50 short prolateral macrosetae; ti II p1-1-0, v2-2-2; mt II p1-0-1, r0-0-1, v2-2-2-2-2-2; ta II v7; fe III d5, dp4, dr1, v11; ti III p1-1-0, vp1-1-0, r1-1-0; mt III p1-0-1, r1-1-1, v2-2-2-2- 2-2, dr-0-0-1; ta III v11; fe IV d8; ti IV vp1-1-0; mt IV p0-0-1, r0-0-1, v2-2-2-2-2; ta IV v15.

Palp ( Fig. 20View FIGURE 20): femur 1.5 times longer than carapace, cymbium about as long as bulb, with dense fringe of setae, longer on ventral face, bulb piriform, simple, sperm duct with 3 coils, embolus with conspicuous, translucent keel short, tip bent.

State of the specimen: good, left palp dissected, left leg I disarticulated from tibia, left leg IV disarticulated from trochanter.

Female (Paratype, MACN-Ar 38366). Carapace brown, stippled with dark brown and with dark brown Vshaped median pattern. Labium, endites and sternum dark brown. Legs dark brown, with longitudinal brown stripes, gradually becoming yellow brown on tips, femora and tibia I and II hirsute, with long setae. Abdomen grayish, light brown.

Total length 13.1. Carapace 4.17 long, 3.65 wide. Clypeus 0.72 long. Eye sizes and interdistances: AME 0.16, PME 0.23, ALE 0.28, PLE 0.23, AME-AME 0.068, PME-PME 0.22. Sternum 2.17 long, 2.02 wide. Palp: femur 2.51, height 1.02, tibia length 1.43, height 0.91. Leg I: fe 5.58, pa 1.85, ti 4.9, mt 4.58, ta 2.73. II: fe 4.39, pa 1.48, ti 3.51, mt 3.56, ta missing. III: fe 3.34, pa 1.45, ti 2.67, mt 2.91, ta 1.71. IV: fe 4.92, pa 1.56, ti 4.22, mt 3.96, ta 2.08. Abdomen: length 9.05, width 6.85.

Palp macrosetae on ventral surface of tibia and tarsus. Leg macrosetae: ti I v2-2; mt I p0-0-2, v2-2-2-2-2; ta I v15; tiI II v2-2; mt II p0-0-2, r0-0-1, v2-2-2-2; fe III d2-0-0; tiI III dr1-1-0; mt III p1-0-1, r1-1-1, v2-2-2-2; ta III v12; fe IV d3-0-0; ti IV vp-1-1-1; mt IV p0-0-1, r0-0-1, v2-2-2-2-2; ta IV v14. Calamistrum with 3 rows with tightly packed and neatly arranged setae, rows with ~20-18-11 setae, prolateral one extending well beyond calamistrum crest.

Endogyne ( Fig. 23View FIGURE 23) without membranous portion, spermathecae digitiform, inner spermathecae with glandular pores throughout, outer spermathecae with glandular pores restricted to apex.

State of the specimen: good, left leg I disarticulated from tibia, right legs II–III set apart for molecular work, genitalia dissected.

Variation. Females (n = 3): total length 12.50–13.10 (12.87), carapace length 4.17–5.03 (4.67), femur I length 5.25–6.22 (5.68), femur/carapace ratio 1.09–1.34 (1.22). The genital morphology of females varies, especially regarding the length of the spermathecae.

Type material. Holotype ³: MOROCCO: Guelmin-Es Semara : Assa-Zag, Zag, dry shrubland on sand dunes, under large rocks, 28.24872°N, 09.33291°E, leg. M. Stockmann, X.2016, 1³ ( SMNH).GoogleMaps 

Paratypes: MOROCCO: Drâa-Tafilalet : Errachidia , Arfoud, Alnif, dry shrubland near mountain ranges, in webs under large rocks, 31.15695°N, 05.04701°E, leg. M. Stockmann, X.2016, 2♀ ( HUJ 20310View Materials)GoogleMaps  , 1♀ (MACN-Ar 38370), 3♀ (MACN-Ar 41204), 1³ ( SMF)  . Guelmin-Es Semara : Assa-Zag , Assa, Touizgui, dry shrubland in wadi, in long webs under large rocks, 28.3836°N, 09.38865°W, leg. M. Stockmann, X.2016, 1♀ (MACN-Ar 38369)GoogleMaps  ; Zag , dry shrubland on sand dunes, under large rocks, 28.24872°N, 9.33291°W, leg. M. Stockmann, X.2016, 2♀ ( HUJ 20312View Materials)GoogleMaps  , 1♀ (MACN-Ar 38366), 3♀ (MACN-Ar 41203), 1♀ ( SMNH)  , 1♀ ( SMF)  , 1³ (MACN-Ar 39465). Rio de Oro : Cape Bojador, 26.12694°N, 14.49917°W, leg. D. Van Damme, 20.VIII.1973, 2♀ 2 imm. ( MRAC 167943View Materials)GoogleMaps  ; 20 miles E Tiznit , 29.68327°N, 9.39203°W, 400 m a.s.l., leg. E.S. Ross, 13.VII.1981, 1♀ ( CAS 9053476View Materials)GoogleMaps  .

Additional material examined. We have examined females closely related to S. wunderlichi  sp. nov., but whose identification is pending until males are discovered; these have not been added to the distribution map.GoogleMaps  CHAD: Massif du Tibesti, 00.77619°N, 18.04932°E, leg. Y. Brandily, VII–X.1965, 1♀ ( MRAC 132960View Materials).GoogleMaps  EGYPT: Ein Suchna: Eastern Desert , dried, salty area with many Tamarisk , 29.63262°N, 32.33009°E, leg. M. Alderweireldt, 20. III.2000, 1♀ ( MRAC 209671View Materials).GoogleMaps  Hurghada: Abu Mun Ahr island , in web in crab hole in mangrove, 27.1853°N, 33.81039°E, leg. RGoogleMaps  . Jocqué & F. Jocqué, 6.IV.1996, 1♀ ( MRAC 203038View Materials). SUDAN: Dongola , 19.16983°N, 30.47489°E, 1♀ ( ZMB 592View Materials). NO COUNTRY DATAGoogleMaps  : Mediterranean region , 1♀ ( ZMB 3281View Materials)  .

Distribution. Morocco ( Fig. 3View FIGURE 3).

Natural history. Near Zag, specimens were found in a flat and wide area surrounded by sand dunes, mostly covered by loose sand but with patches of loamy soil and sparse shrubs and small trees; Sahastata  were found under large rocks. In Alnif, specimens were taken in a similar habitat, but with loamy rather than sandy soil. North of Zag, specimens were collected in a wadi with more rocky and loamy soil, including more shrubs, plants and small trees than the first locality. Specimens seemed to occupy old burrows of lizards or scorpions beneath boulders to build long, tube-shaped webs, and some females had egg sacs.


Department of Paleozoology, Swedish Museum of Natural History


Hebrew University


Forschungsinstitut und Natur-Museum Senckenberg


Departamento de Geologia, Universidad de Chile














Sahastata wunderlichi

Magalhaes, Ivan L. F., Stockmann, Mark, Marusik, Yuri M. & Zonstein, Sergei L. 2020

Sahastata nigra Benoit, 1968: 97

Marusik, Y. M. & Zamani, A. & Mirshamsi, O. 2014: 9
Benoit, P. L. G. 1968: 97