Fortiseode pervalimand Jia & Ren

Jia, Ting, Liang, Hongbing, Chang, Huali & Ren, Dong, 2011, A new genus and species of fossil Eodromeinae from the Yixian Formation of Western Liaoning, China (Coleoptera: Adephaga: Trachypachidae), Zootaxa 2736, pp. 63-68 : 66-68

Fortiseode pervalimand Jia & Ren sp. nov.

( Figs 1 , 2 A–B)

Etymology. The specific name of pervalida is derived from the Greek word of “ pervalid -” (meaning strong), referring to the robust mandibles.

Material. CNU-COL-LB-2010495, deposited in the Key Lab of Insect Evolution & Environmental Changes, Capital Normal University, Beijing, China.

Locality and horizon. Collected from 2nd bed of Yixian Formation in Huangbanjigou, near Chaomidian Village, Shangyuan County, Beipiao City, Liaoning Province, China; Late Jurassic-Early Cretaceous.

Diagnosis. Same as the genus.

Description. Body small sized, length 10 mm, width 3.5 mm; elytron 5 mm long, with 8 longitudinal striae. The holotype is a dorso-ventral compression of a carabid. Antennae unisetose, scape long and robust, not completely visible from dorsal side; elytra without large pores.

Head. Head sub-triangular, convex; narrower than prothorax; eyes moderately prominent; antennae inserted between eyes and base of mandibles under a frontal ridge, filiform, with eleven antennomere; antennomere 1 (scape) long and robust, antennomere 2 shorter than scape, antennomeres 6 to 10 subequal in length, apical antennomere narrower and longer; mandible falciform, robust, prominent, apices rather blunt, laterally with setae, occlusal margins toothed, length equal to that of head, molar plate visble, outer sides of mandibles not expanded; base of head retracted into pronotum.

Thorax. Prothorax width nearly equal to that of paired elytra together, with lateral margins inflexed and a distinct submarginal suture between proepipleuron and propleuron; pronotum cordiform, length 2mm, width 3.4mm, widest at about anterior one-fifth of the pronotum, anterior edge of pronotum strongly concave, with width 3.1mm, posterior edge straight, width 2.1mm. base of pronotum with similar width as base of elytra; pronotum with lateral margins narrowly reflexed; forecoxal cavities open, confluent, bridged; midcoxal cavities disjunct, confluent; hind coxal cavities incomplete, confluent.

Pterothorax. Scutellum triangular, visible; elytra margined laterally and basally, humeri broadly rectangular; posteriorly not sinuate, apical margin narrowly rounded, elytra broadest in the middle, with basal border, width 2.5 mm, wider than that of posterior edge of pronotum, length 6 mm, twice as long as wide, dorsal surface striate, with 8 longitudinal striae, each stria linear, striae absent laterally, surface smooth, without large pores; second elytral interval with similar width as other intervals, elytra margin with internal plica toward apex.

Abdomen. Elytra completely cover the last ventrite of abdomen, abdomen with six sterna (2–7), sternum 2 (first visible sternum) interrupted by hind coxae, remnants visible only at sides; the last four ventrites of abdomen are much shorter compared with the first two ventrites; the first abdominal segment with length 0.7 mm, the second segment longest, with length 0.9 mm, segments 3 to 5 equal in length 0.3 mm, and the last segment length 0.6 mm.

Note“-” indicate that it is not sure about the age.

Legs. Gressorial, fairly short and thick; legs with strong femora and tibiae, metatarsus shoter than 1/2 of elytra, metatarsus short, with length nearly 1/2 of metatarsus; trochanters large, longer than 1/3 metafemur length, tarsi with 5 segments; tarsomere 5 terminated by pair of claws; claws with inner margins smooth; metatarsi length 2.3 mm, shorter than half of elytra, length 2.3 mm, shorter than half of elytra, nearly equal to half of metatibia; front and middle coxae globular, hind coxae dilated internally, and extended each side to lateral margins of body; metacoxae larger and transverse; metafemur much wider than metatibia; front tibia with large apical spurs, inner margin simple; hind trochanters large, longer than one-third of metafemur.

Figures

FIGURE 1. Fortiseode pervalimand Jia & Ren gen. et sp. nov, holotype. CNU-COL-LB- 2010495.

FIGURE 2. Fortiseode pervalimand Jia & Ren gen. et sp. nov, line drawings of holotype. A. Dorsal view of body; B. Ventral view of body.