Mastigoproctus floridanus Lönnberg, 1897

Barrales-Alcalá, Diego, 2018, Systematic Revision Of The Giant Vinegaroons Of The Mastigoproctus Giganteus Complex (Thelyphonida: Thelyphonidae) Of North America, Bulletin of the American Museum of Natural History 2018 (418), pp. 1-64 : 25

Mastigoproctus floridanus Lönnberg, 1897 ,

stat. nov.

Mastigoproctus giganteus floridanus Lönnberg, 1897: 189–191 (synonymized by Kraepelin, 1899: 224); McMonigle, 2013: fig. unnumbered, 100 (part).

Mastigoproctus giganteus: Kraepelin, 1899: 224 ; Banks, 1900: 422 (part); 1904: 143; Howard, 1919: 26; Ewing, 1928: 41–43, fig. 1; Savory, 1964: 166 (part); Muma, 1967: 21–23, fig. 16; Weygoldt, 1970: 1–7, figs. 1a, b, 2a–d, 3a–f, 4a–e; 1971: 137–141, figs. 1a, b, 2a, b, 3, 4, 5a–c, 6, pl. Ia–d, IIa–c, IIIa–c; Browder, 1973: 161; Jespersen, 1978: 241, figs. 21–23; Coddington et al., 1990: 11 (part); Corey and Stout, 1990: 169, 170; Crawford, 1990: 434, fig. 16.5 (part); Folkerts et al., 1993: 161; Punzo, 2001: 35–39, figs. 3, 4; 2005a: 684–690; 2005b: 172, 173; Beccaloni, 2009: 111 (part); Carrel and Britt, 2009: 500–502, figs. 1–4 (part); Kern and Mitchell, 2011: 2, 4, figs. 1–5 (part).

TYPE MATERIAL: U.S.A.: Florida: holotype ♀? (Swedish Museum of Natural History, Stockholm) [not examined].

DIAGNOSIS: Mastigoproctus floridanus most closely resembles M. giganteus , from which it differs as follows. In the adult male of M. floridanus , spines S4 and S5 on the prodorsal margin of the pedipalp trochanter are blunt and a short, blunt subapical accessory spine ( AS) is present whereas, in the adult male of M. giganteus , all spines on the prodorsal margin of the pedipalp trochanter are sharply pointed and an AS is absent.

Mastigoproctus floridanus also differs from M. cinteotl , sp. nov., and M. tohono , sp. nov., as follows. The carapace epistoma is visible in dorsal aspect in M. floridanus but not in M. cinteotl and M. tohono , and the lateral borders of the carapace are almost smooth in M. floridanus but strongly serrate in M. tohono . Spines S4 and S5 on the prodorsal margin of the pedipalp trochanter of the adult male are short and blunt in M. floridanus whereas S4 and S5 are longer and sharply pointed in M. cinteotl , and S4 is shorter in M. tohono .

DESCRIPTION: The following description is based on three males and two females.

Total length: Maximum length from anterior margin of carapace to posterior margin of opisthosomal segment XII (pygidium) in adult specimens 51.8 mm (♂) and 60.6 mm (♀) (table 2).

Color: Carapace dark reddish brown, anterior and lateral margins dark brown. Tergites I –IX dark reddish brown (♂) to reddish brown (♀), posterior and lateral borders darker. Sternites I and II light reddish brown mesally, margins reddish brown ; III –IX and pygidium reddish brown (♀) to dark reddish brown (♂). Flagellum light reddish brown, segments with reddish-brown macrosetae. Pedipalp trochanter, femur, patella, and tibia dark reddish brown, mesal surfaces with few reddish macrosetae ; patellar apophysis, fixed finger, and basitarsus blackish, with few reddish macrosetae. Legs I– IV reddish brown ; II–IV coxae, ventral surfaces light reddish brown.

Carapace: Epistoma visible in dorsal aspect. Pair of strongly developed carinae anterolaterally, extending from lateral ocelli to two-thirds the distance to median ocular tubercle (fig. 10F). Three pairs of lateral ocelli with accessory translucent eyespot situated between anterior and posterior ocelli; anterior ocelli larger than median and posterior ocelli; anterior two pairs slightly separated from posterior pair by tubercle, distance between median and posterior ocelli 3× distance between anterior and median ocelli. Median ocular tubercle smooth, situated in anterior 10% of carapace (table 2); distance between ocelli almost 2× ocular diameter. Carapace sur-