Nemoleontini Banks, 1911

Suryanarayanan, Thangalazhi Balakrishnan, Bijoy, Chenthamarakshan & Ábrahám, Levente, 2022, Redescription of Banyutus cubitalis (Navás, 1914) (Neuroptera, Myrmeleontidae) and key to antlion genera in tribe Nemoleontini from India, Zootaxa 5182 (1), pp. 64-74 : 66

Key for the genera of the tribe Nemoleontini Banks, 1911 known from India ( Fig. 2 )

1. A 2 and A 3 unit in short term then diverge, A 2 simple, A 3 forked in FW ( Fig. 2 A ), male gonocoxites 9 fused as a forked.... 2

- A 2 and A 3 unit in short term then diverge, A 2 forked, A 3 simple in FW ( Fig. 2 B ), male gonocoxites 9 shaped as a pair of plates............................................................................................. 10

2. On basal half of FW Cua (also Cua 1, Cua 2) long and parallel with hind margin (( Fig. 2 C )............................ 3

- On basal half of FW Cua 2 short and not parallel with hind margin ( Fig. 2 A )....................................... 4

3. Tibial spurs long and straight as long as elongated tarsal segment 1, tarsal claws with prominence ( Fig. 2 D and E )................................................................................. Pseudoformicoleo van der Weele*

- Tarsal segment 1 short, tibial spurs as long as tarsi 1-4 combined, tarsal claws simple ( Fig. 2 F and G ).... Creoleon Tillyard

4. Tibial spurs considerably shorter than tarsal segment 1, male ectoproct with short processes ( Fig. 2 H )..... Delfimeus Navás

- Tibial spurs considerably longer than tarsal segment 1, male ectoproct with long distal processes ( Fig. 2 I )............... 5

5. Cup in FW very short runs to A1 directly ( Fig. 2 J )........................................................... 6

- Cup in FW partly parallel before connecting with A1 ( Fig. 2 K )................................................. 7

6. Smaller species (FW 15-20 mm), pronotum V or U-shaped pattern............................ Geyria Esben-Peterson

- Larger species (FW longer than 20 mm), pronotum with two lateral stripes....................... Macronemurus Costa

7. Wings narrow, both margins almost parallel, tarsal segment 1 and tibial spurs elongated as long as or longer than segment 2-4 combined, male abdomen usually very long ( Fig. 2 L )........................................... Nemoleon Navás

- Wings long oval, tarsal segment 1 not elongated, tibial spurs varied in length..................................... .. 8

8. Smaller species (FW shorter than 30 mm in India), tibial spurs shorter than tarsal segment 1-4 together or missing............................................................................................... Neuroleon Navás

- Larger species (FW longer than 30 in India), tibial spurs almost as long as tarsal segment 1-4 together.................. 9

9. Tarsal claws enlarged, at least three fourth part of tarsal segment 5, tibial spurs slightly curved, posterior Banksian line slightly visible in FW ( Fig. 4 C ).................................................................... Banyutus Navás

- Tarsal claws reach only half length of tarsal segment 5, tibial spurs obviously curved and strong, Banksian line not visible in FW................................................................................... Distoleon Banks

10. Tibial spurs as long as tarsal segment 1, claws opposable.............. Paraglenurus van der Weele / Negrokus Navás **

- Tibial spurs as long as tarsal segment 1-2 combined, claws not opposable......... Indoleon Banks / Indophanes Banks ***

Figures

FIGURE 2. Diagnostic characters for the Indian Nemoleontini tribe, red arrows show the characters mentioned in the key A— Distoleon sp. basal part of FW; B—Paraglenurus sp. basal part of FW; C—Creoleon sp. FW; D and F –Pseudoformicoleo sp. tibial spurs and claws; E and G—Creoleon sp. tibial spurs and claws; H—Delfimeus sp.male ectoproct; I and J—Macronemurus sp. male ectoproct; K and L—Nemoloen sp. male habitus.

FIGURE 4. Banyutus cubitalis (Navás, 1914). A—head in frontal view; B—head and thorax in dorsal view, C—male wing venation, D—foreleg; E—hindleg in lateral view.