Madecorphnus

Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, Zootaxa 4207 (1), pp. 1-93 : 23-24

Key to the Madecorphnus species (males)

1. Right mandible with a tooth on inner side of scissorial part ( Fig. 9 A).............................................2

- Right mandible without a tooth on inner side of scissorial part ( Fig. 13 A).........................................5

2. Mentum without tubercles; parameres slender in dorsal view ( Fig. 9 B); internal sac of aedeagus with 2 large bifurcated scler- ites and a few dozen spinules ( Figs. 9 E–F)................................................ M. falculoides (Paulian)

- Mentum with 2 tubercles ( Fig. 11 F); parameres and internal sac not as above..................................... 3

3. Internal sac of aedeagus with 3 sclerites, 2 larger and 1 smaller ( Figs. 10 E–F)...................... M. montreuili Frolov

- Internal sac of aedeagus with 2 bifurcated sclerites ( Figs. 11 D–E, 12C)...........................................4

4. Sclerites of internal sac of aedeagus smaller ( Figs. 11 D–E); body narrower and eyes smaller ( Fig. 11 A)... M. barclayi Frolov

- Sclerites of internal sac of aedeagus larger ( Fig. 12 C); body wider and eyes larger ( Fig. 12 A)....... M. tuberculatus Frolov

5. Internal sac of aedeagus with long sclerites but without fields of small spinules ( Fig. 13 D)............... M. fisheri Frolov

- Internal sac armature not as above....................................................................... 6

6. Apices of parameres without lateral teeth ( Fig. 14 C)..........................................................7

- Apices of parameres with more-or-less developed lateral teeth ( Fig. 16 C).........................................8

7. Parameres without lateral excavations, about 1.5 times shorter than basal sclerite of aedeagus ( Fig. 14 C).. M. pauliani Frolov

- Parameres with lateral excavations, about 2 times shorter than basal sclerite of aedeagus ( Fig. 15 B)........ M. niger Frolov

8. Head and pronotum with fine, dense punctation; internal sac of aedeagus with 3 large sclerites and an area with smaller spinules ( Figs. 16 E, F)................................................................... M. punctatus Frolov

- Head and pronotum with indistinct to sparse punctation (punctures separated by at least 3 times their diameter); internal sac of aedeagus with different armature.........................................................................9

9. Parameres with large lateral teeth ( Fig. 17 B)...............................................................10

- Parameres with small lateral teeth ( Fig. 19 B)............................................................... 11

10. Internal sac of aedeagus with 1 long, spinule-shaped sclerite and with 3 small, short spinules ( Figs. 17 D–E).................................................................................................... M. dentatus Frolov

- Internal sac of aedeagus with 2 tooth-shaped sclerites, 1 longer sclerite, and a few fields of minute spinules, 1 of which forms a finger-shaped process of the sac ( Fig.18 C)............................................ M. saintemariensis Frolov

11. Internal sac of aedeagus with 1 sclerite composed of 2 long slender spinules connected basally ( Figs. 19 D–E)................................................................................................... M. simplex Frolov

- Internal sac of aedeagus with 2 or more separate sclerites.....................................................12

12. Internal sac of aedeagus with 2 rather large sclerites: a spur-like sclerite, and an asymmetrical, claw-shaped sclerite ( Figs. 20 D–E)............................................................................. M. aquilonius Frolov

- Internal sac of aedeagus with different armature............................................................13

13. Parameres with small teeth at extreme apex (Fig. 21B, lateral view); body length greater than 7.0 mm.... M. falcatus Paulian

- Parameres of different shape; body length less than 6.0 mm...................................................14

14. Internal sac of the aedeagus with 2 large, curved paired sclerites and 1 long, spur-shaped sclerite ( Figs. 22 D–E).…………. 15

-. Internal sac armature different.......................................................................... 16

15 Curved paired sclerites larger, semicircular; spur-shaped sclerite shorter........................ M. falciger (Lansberge)

- Curved paired sclerites smaller, comma-shaped; spur-shaped sclerite longer.......................... M. hanskii Frolov

16 Internal sac of aedeagus with 2 large, separate sclerites and numerous small spinules, some of which are robust and almost tooth-shaped and may be arranged into 2 indistinctly separated clusters.......................... M. cuccodoroi Frolov

- Internal sac of aedeagus with 2 or more separate sclerites and numerous small spinules; the latter are fine and are all approxi- mately the same size..................................................................................17

17 Parameres less acute in lateral view ( Fig. 25 B); internal sac of the aedeagus with 1 longer sclerite and 1 smaller, somewhat bifurcated sclerite ( Figs. 25 D–E).......................................................... M. brunneus Frolov

- Parameres more acute in lateral view ( Fig. 26 B); internal sac with 2 longer (1 doubled) and 2 smaller sclerites ( Figs. 26 D–E)................................................................................................... 18

18 Parameres less tapering apically, lateral teeth closer to paramere apices ( Fig. 26 B); eyes larger, rounded in dorsal view ( Fig. 26 A)............................................................................... M. perinetensis Frolov

- Parameres more tapering apically, lateral teeth not so close to paramere apices ( Fig. 27 B); eyes smaller, narrowly elongate in dorsal view ( Fig. 27 A)................................................................. M. peyrierasi Frolov

Figures

FIGURE 9. Madecorphnus falculoides. Habitus in dorsal view (A—male, C—female), aedeagus in lateral view and parameres in dorsal view (B), labels of holotype (D), internal sac of aedeagus (E—photograph, F—schematic representation of armature), distributional records (G).

FIGURE 10. Madecorphnus montreuili. Habitus in dorsal view (A—male, C—female), aedeagus in lateral view and parameres in dorsal view (B), labels of holotype (D), internal sac of aedeagus (E—photograph, F—schematic representation of armature), distributional records (G), protibia (H, male left, female right).

FIGURE 11. Madecorphnus barclayi. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels of holotype (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), mentum (F), distributional records (G).

FIGURE 12. Madecorphnus tuberculatus. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), internal sac of aedeagus (C), distributional records (D).

FIGURE 13. Madecorphnus fisheri, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D), distributional records (E).

FIGURE 14. Madecorphnus pauliani, holotype. Habitus in dorsal view (A), labels (B), aedeagus in lateral view and parameres in dorsal view (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).

FIGURE 15. Madecorphnus niger. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), internal sac of aedeagus (C—photograph, D—schematic representation of armature), distributional records (E), labels of holotype (F).

FIGURE 16. Madecorphnus punctatus, holotype. Habitus (A) and head (B) in dorsal view, aedeagus in lateral view and parameres in dorsal view (C), labels (D), internal sac of aedeagus (E—photograph, F—schematic representation of armature), distributional records (G).

FIGURE 17. Madecorphnus dentatus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).

FIGURE 18. Madecorphnus saintemariensis, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), internal sac of aedeagus (C), distributional records (D).

FIGURE 19. Madecorphnus simplex, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).

FIGURE 20. Madecorphnus aquilonius, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).

FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature).

FIGURE 25. Madecorphnus brunneus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).

FIGURE 26. Madecorphnus perinetensis, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).

FIGURE 27. Madecorphnus peyrierasi, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).