Triodontus

Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, Zootaxa 4207 (1), pp. 1-93 : 58

publication ID	https://doi.org/ 10.11646/zootaxa.4207.1.1
publication LSID	lsid:zoobank.org:pub:F2875582-31E2-496F-AEEF-1D657DD86C33
DOI	https://doi.org/10.5281/zenodo.6057680
persistent identifier	https://treatment.plazi.org/id/5B4FBD00-FFE8-6153-FF02-63E4FEE3F95C
treatment provided by	Plazi
scientific name	Triodontus
status

Treatment

Key to Triodontus species (males)

1 Central pronotal bulge with 3 more-or-less similar tubercles ( Figs. 3 1 View FIGURE 3 A–B); outer lobes of parameres rather large, mostly covering inner lobes (in lateral view: Fig. 31 View FIGURE 31 E).............................................................. 2

- Central pronotal bulge with 2 tubercles ( Fig. 35 View FIGURE 35 A); outer lobes of parameres smaller, not covering inner lobes (in lateral view: Fig. 35 View FIGURE 35 E).......................................................................................... 4

2 Body length less than 9.0 mm........................................................................... 3

- Body length 9.5–12.0 mm................................................................ T. hova (Fairmaire)

3 Abdominal sternite 8 narrower and more sinuate medially, without oval transverse concavity ( Fig. 31 View FIGURE 31 F); lateral lobes of parameres with small, keel-shaped processes on proximal parts ( Fig. 31 View FIGURE 31 E)................. T. nitidulus (Guérin-Méneville)

- Abdominal sternite 8 wider and less sinuate medially, with oval transverse concavity ( Fig. 33 View FIGURE 33 G); lateral lobes of parameres without small, keel-shaped processes on proximal parts ( Fig. 33 View FIGURE 33 F)............................. T. bicavatus (Fairmaire)

4 Abdominal sternite 8 with more-or-less developed tubercule medially ( Fig. 34 View FIGURE 34 F)................................... 5

- Abdominal sternite 8 without tubercule medially ( Fig. 36 View FIGURE 36 G)............................................. ………. 6

5 Tubercules of abdominal sternite 8 rounded apically ( Fig. 34 View FIGURE 34 F); lateral lobes of parameres with keel-shaped processes on prox- imal parts (in dorsal view, Fig. 34 View FIGURE 34 D).............................. T. ankarafantsikae Frolov, Montreuil & Akhmetova

- Tubercule of abdominal sternite 8 with concavity apically ( Fig. 35 View FIGURE 35 G); lateral lobes of parameres with semicircular notches on proximal parts (in dorsal view, Fig. 35 View FIGURE 35 E)................................. T. lemoulti Frolov, Montreuil & Akhmetova

6 Body length less than 6.5 mm ......................................................... T. modestus (Benderitter) View in CoL

- Body length 9.0–12.0 mm.............................................................................. 7

7 Outer lobes of parameres strongly sclerotized, with distinct lateral notches and teeth situated more-or-less apically ( Fig. 3 7 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 D)................................................................................................... 8

- Outer lobes of parameres less sclerotized, somewhat rolled up in most species; lateral notches, if present, situated more basally ( Fig. 41 View FIGURE 41 E)......................................................................................... 10

8 Outer lobes of parameres slender in lateral and dorsal view ( Fig. 37 View FIGURE 37 D)............................. T. owas Westwood

- Outer lobes of parameres wider, at lease in dorsal view ( Figs. 39 View FIGURE 39 E, 40E)......................................... 9

9 Outer lobes of parameres with slit-shaped lateral notches ( Fig. 39 View FIGURE 39 E)............ T. viettei Frolov, Montreuil & Akhmetova

- Outer lobes of parameres with wide lateral notches ( Fig. 40 View FIGURE 40 E)............................ T. hildebrandtii (Fairmaire)

10 Outer lobes of parameres long, curved downward and inward, with more-or-less separated strongly sclerotized lateral parts and membranous, overlapping medial parts; lateral notches absent ( Fig. 41 View FIGURE 41 E)........................... T. alticola Paulian

- Outer lobes of parameres of different shape, without distinctly separated lateral and medial, membranous parts; lateral notches absent or present..................................................................................... 11

11 Outer lobes of parameres without lateral notches ( Fig. 42 View FIGURE 42 F)................ T. fairmairei Frolov, Montreuil & Akhmetova

- Outer lobes of parameres with more-or-less distinct lateral notches and teeth ( Figs. 43 View FIGURE 43 F, 44D, 45E, 46D).............. 12

12 Outer lobes of parameres with distinct, slit-shaped lateral notches (in lateral view, Figs. 43 View FIGURE 43 F, 44D)....................13

- Outer lobes of parameres less distinct, wider lateral notches (in lateral view, Figs. 45 View FIGURE 45 E, 46D)......................... 14

13 Abdominal sternite 8 without fossa ( Fig. 43 View FIGURE 43 E); slit-shaped lateral notches of outer lobes of parameres longer ( Fig. 43 View FIGURE 43 F).................................................................................................. T. itremoi

- Abdominal sternite 8 with shallow but distinct, somewhat transversal fossa ( Fig. 44 View FIGURE 44 E); slit-shaped lateral notches of outer lobes of parameres shorter ( Fig. 44 View FIGURE 44 D)..................................................... T. copridoides Paulian

14 Outer lobes of parameres 1.3 times longer than inner lobes; inner lobes smaller (in lateral view) ( Fig. 45 View FIGURE 45 E).. T. hanskii Frolov

- Outer lobes of parameres as long as inner lobes; inner lobes larger (in lateral view, Fig. 46 View FIGURE 46 D) T. inexpectatus Frolov, Montreuil & Akhmetova