Poecilosomella subpilimana, Papp, 2002

Papp, L., 2002, Eighteen New Oriental Species Of Poecilosomella Duda (Diptera: Sphaeroceridae), Acta Zoologica Academiae Scientiarum Hungaricae 48 (2), pp. 107-156 : 145-156

Poecilosomella subpilimana sp. n.

( Figs 65–68 )

Holotype male ( ROM): INDONESIA, Sumatra, W. Sumatra: Kerinci-Seblat Nat. Pk., trial to

L. Tujuh, 14 SEP 1989. B Hubley, DC Darling, ROM 893074 – 1° rainforest, 1640 m, screen sweep,

1°42’N, 101°19’E. Paratype male ( HNHM): same data as for the holotype .

Measurements in mm: body length 2.65 (holotype), 2.45 (paratype male), wing length 2.48 (holotype), 2.42 (paratype), wing width 1.24 (holotype), and 1.16 (paratype).

Frons dull greyish red, facial plate shiny, face, cheeks and genae reddish. Two pairs of strong orbital setae close to each other. Two pairs of short but not thin ifr. Genal seta only 0.125 mm long. Scape brown, pedicel yellow, first flagellomere reddish grey, covered with very dense white cilia. Pedicel is somewhat longer than first flagellomere (0.16 mm vs. 0.15 mm). Arista with short cilia only.

Mesonotum dull black with 4 sagittal, 4 pairs of dorsocentral and 3 pairs of lateral silvery spots. Anterior katepisternal thin and short. Scutellum as long as wide basally, concolorous with the mesonotum, with 6 silvery spots: 2 sagittal (basal and medial), 2 larger spots basolaterally including the bases of lateral scutellar setae and a pair of smaller spots on bases of apical scutellars.

Wing almost unicolorous light brown, veins yellow, only cross-veins lighter, one brown spot subapically on R 1, which also covers Rs fork, another spot on costa and wing plane proximally on second costal section and a third one around the upcurving section of R 2+3. Costal hairs short and less dense than in the borboroides group. Second costal section as long as third section. Vein R 2+3 without a vein appendage subapically. Apical half of vein R 4+5 straight, medial part more strongly curved.

All the femora and tibiae slightly thickened. Legs mainly dark brown but apical half of femora dorsally lighter (maybe the type specimens were kept in alcohol before pinning), tibiae with broad light ring medially and subapically. No parts of fore tarsomeres white but apical part of basitarsus and of the 2nd tarsomere light yellow, 3rd tarsomere wholly light yellow on all three tarsi. Male fore tibia and basitarsus with hairs much longer than diameter of tibia/basitarsus at their bases. Male mid tibia not curved inwards and not flattened, ventrally with shorter hairs (only some posterior hairs longer than tibial diameter). Mid tibia without a true ventroapical but with three, short, thick black setae instead. Mid tibial setae: no strong anterior or ventral seta, anterodorsals at 1/5 (short), at 3/10 (longer) at 11/15 (short) and at 5/6 (strong), posterodorsals (all strong) at 3/10, 3/5 and 5/6.

Male abdominal sternite 5 ( Fig. 65 ) asymmetrical, medial part broadly emarginate, area between sternite 5 & 6 with long fine hairs arranged in 2 rows. Sternite 6 with a weakly sclerotized medial plate, which bears, asymmetrically, fine hairs as well as minute thornlets. Epandrium short with = postgonite, lateral view. Scales: 0.2 mm for Fig. 65–66 , 0.1 mm for Figs 67–68

short setae. Cerci normal. Subepandrial sclerite ( Fig. 66 ) rather simple with a sagittal furrow, covered partly by black thornlets and with some medium long setae not only marginally. Male surstylus ( Fig. 67 ) bilobed: anterior lobe incised apically, with numerous long setae, posterior lobe enlarged caudally with a strong apical tooth and subbasally with an extremely long flat thorn-like process. This process is curved mediocaudally, following the curvature of surstyli and epandrium, and is bordered by long, but slightly curved, setae. Postgonite ( Fig. 68 ) rather simple: twice curved but medial part almost straight, with blunt apex and with a limited number of minute posterior hairlets.

Female unknown.

Distribution: Indonesia (Sumatra).

P. subpilimana sp. n. is not an easily recognisable species by body characteristics. It is hypothesized that this species is not a member of the species-group of P. himalayensis , P. annulitibia and P. nepalensis , centred in distribution in the Himalayas (see DEEMING 1969), although it keys with himalayensis on account of its similar wing venation and male mid tibial structures. The only reliable characters for its identification are in the male genitalia; in this respect it is similar to P. pilimana sp. n., particularly because of the extremely long black process (spine) on its surstylus.

A KEY TO THE ORIENTAL SPECIES OF POECILOSOMELLA (based on PAPP 1991 a)

1 (14) Vein R 2+3 with a distinct vein appendage (stump vein) subapically, i.e. at the turning point of curvature to costa.

2 (9) Second costal section much shorter than third. Male mid tibia and basitarsus without long cilia. No mid ventral seta on mid tibia.

3 (4) No ventroapical on mid tibia, 3 thick, short setae instead. A long upcurved genal bristle present. Male fore tibia and 2 basal tarsomeres with long dense hairs on whole of ventral half. Surstylus very characteristic ( PAPP 1991 a:

fig. 18), laterally with a long, curved, blade-like thorn. Postgonite ( PAPP

1991 a: fig. 20) much curved apically. Taiwan, Viet Nam

P. furcata ( DUDA, 1925)

4 (3) Mid tibia with a distinct ventro-apical seta.

5 (6) Wing with a diffuse dark brown spot on both sides of vein R 4+5, i.e. in middle of the r 4+5 radial and medial cells. A long upcurved genal bristle present, much longer than genal diameter below eye, present. Male fore tibia and 2 basal tarsomeres with long dense hairs on whole of ventral half. Sabah P. conspicua sp. n.

6 (5) Wing evenly brown, without diffuse brown spots in middle of the r 4+5 radial and medial cells. Genal bristle weak, shorter than genal diameter below eye. Male fore tibia and basal tarsomeres without long dense hairs.

7 (8) Ventroapical seta on mid tibia slightly curved and shorter than diameter of tibia subapically. Male surstylus ( Fig. 14 ), postgonite ( Fig. 15 ). Java, Sumatra P. meijerei ( DUDA, 1925)

and 6, ventral view. Scale: 0.2 mm for Fig. 72 , 0.1 mm for Figs 69–71

8 (7) Ventroapical seta of mid tibia much longer than diameter of tibia subapically. Taiwan P. ronkayi sp. n.

9 (2) Second costal section about as long as third. Male mid tibia and basitarsus with or without long cilia. Mid tibia with or without mid ventral seta. No strong upcurving genal bristle.

10 (11) Mid tibia without an anterior seta, or a small anterior close to the distal anterodorsal. Male mid tibia with a very long ventral spur (half as long as basitarsus). Female first flagellomere enlarged (much longer than scape and pedicel combined), with long cilia apically. Taiwan

P. longicalcar sp. n., pro parte

11 (10) Mid tibia with a strong anterior seta. Male mid tibia with a short ventroapical seta, or ventroapical indiscernible. Female first flagellomere normal.

12 (13) Anterior bristle of mid tibia equidistant between the two strong anterodorsals. Male subepandrial sclerite with a blunt shovel-formed appendage. Postgonites arcuately curved and pointed. Nepal, Taiwan

P. annulitibia ( DEEMING, 1969)

13 (12) Anterior bristle of mid tibia much nearer to proximal anterodorsal than to long distal anterodorsal. Male subepandrial sclerite with a long medial process. Postgonites slightly curved and blunt apically. Nepal, India, Pakistan P. nepalensis ( DEEMING, 1969)

14 (1) Vein R 2+3 without a vein appendage subapically.

15 (24) Fore tarsomeres, and sometimes fore basitarsus partly, clear white. Second costal section much shorter than third.

16 (21) Costa terminates at vein R 4+5. Wings completely dark. Arista with long cilia (similar to that of borboroides and ornata ) or genal bristle very small.

17 (18) Hind tibia all dark, without any light ring. Mid tibia with a long ventroapical seta (much longer than tibial diameter subapically). Genal seta very small. No ventral seta on the middle of mid tibia. Philippines, Sri Lanka

P. albipes ( DUDA, 1925)

18 (17) Hind tibia with one yellow ring medially and one apically. No ventroapical on mid tibia or ventroapical seta much shorter than tibial diameter subapically. Genal seta normal.

19 (20) Two pairs of thin but subequal orbital setae. No mid ventral seta on mid tibia. Second costal section only slightly longer than half third section. Vein R 2+3 meets costa at an acute angle. Fore tarsomeres 2–5, mid and hind tarsomeres 3–5 clear white. Philippines P. hayashii sp. n.

20 (19) Anterior orbital seta reduced to a minute hair (just discernible), posterior orbital setae long and thick. Mid tibia with a strong mid ventral seta. Second costal section about 3/5 length of third section. Vein R 2+3 meets costa perpendicularly. Fore tarsomere 5, as well as mid and hind tarsomeres 4–5 all dark. Sumatra P. brevisecunda sp. n.

21 (16) Costa terminates distinctly beyond tipof vein R 4+5. Wings lighter with dark spots (distinct pattern). No ventroapical on mid tibia or ventroapical seta much shorter than tibial diameter subapically.

22 (23) Costa terminates not far from apex of R 4+5. Male mid tibia with very long hairs and with straight stiff setae ( DEEMING 1969: fig. 3). Nepal, India, Sri Lanka, Indonesia (W. Flores, Lombok) P. aciculata ( DEEMING, 1969)

23 (22) Costa overruns apex of R 4+5 by a distance of the length of dM-Cu crossvein. Male mid tibia ventrally with a short ventro-apical but with short hairs only (shorter than tibial diameter). Sumatra

P. paraciculata sp. n.

24 (15) No parts of fore tarsomeres white, although they are at least partly yellow.

25 (34) Arista with very long cilia. Costal hairs rather long. Second costal section much shorter than third. Male mid tibia and basitarsus ventrally without long hairs, or tibia with a limited number of long hairs.

26 (29) Mid tibia with a strong (antero)ventral bristle at apical 3/5 to 2/3 but no ventro-apical seta on mid tibia.

27 (28) Male surstylus ( Fig. 12 ) with a large tooth and with a process caudally to it. Male subepandrial sclerite thinner digitiform without thicker setae ( Fig.

11). Wing without a diffuse dark spot distally to apex of R 2+3. Widespread

P. ornata (DE MEIJERE, 1908) , sp. restit.

28 (27) Male surstylus with a small tooth and no process behind it ( Fig. 70 ). Male subepandrial sclerite not curved, edged, broader with some thicker setae ( Fig. 69 ). Wing with a diffuse dark spot distally to apex of R 2+3. Philippines P. picturata ( MALLOCH, 1913), sp. restit.

29 (26) No ventral or anteroventral setae on mid tibia, except for the ventral seta at the apex.

30 (31) Antennae blackish. Wing with a diffuse dark spot distally to apex of R 2+3, male surstylus ( Fig. 21 ) with a single tooth, i.e. without a row of thick black teeth. Taiwan P. formosana sp. n.

31 (30) Antennae reddish or yellowish. Wing without a diffuse dark spot distally to apex of R 2+3, male surstylus ( Figs 4–5 , 7 ) with a row of 4 or 6 thick black teeth.

32 (33) Male surstylus with 6 thick black teeth, ventral process of subepandrial sclerite ( Figs 4–5 ) very broad, postgonite ( Fig. 3 ) less broad. Widespread P. borboroides ( WALKER, 1860)

33 (32) Male surstylus with 4 thick black teeth, ventral process of the subepandrial sclerite ( Fig. 7 ) less broad, postgonite ( Fig. 8 ) very broad. Philippines P. borborus sp. n.

34 (25) Arista with short cilia only. Costal hairs shorter. Second costal section usually as long as, or longer than third section.

35 (38) Scutellum very long, as long as apical scutellar setae or nearly so. Costal ratio mg 2/ mg 3 about 1.5 or even more. All posterodorsals on mid tibia short, i.e. no long paired setae on dorsal half of mid tibia. Male mid tibia with some short curved/inclinate ventroapical spurs.

36 (37) Body reddish, i.e. frons, face and genae red or reddish ochre, at least pleurae partly red. Four (rarely 3) pairs of short and thin ifr. Genae very broad, wings long. No upcurved genal bristle. Female cerci with 2 pairs of very long bent hairs. Nepal, India, Burma, China, Taiwan

P. longinervis ( DUDA, 1925)

37 (36) Body dark, e.g. mesonotum blackish brown. Only 2 or 3 pairs of short and thin ifr. Genae narrower than in longinervis , wings comparatively shorter.

An upcurved genal bristle present. Female cerci with 1 short and 1 moder-

ately long pairs of bristles only. India, Indonesia, Taiwan, Philippines

P. multipunctata ( DUDA, 1925)

38 (35) Scutellum normal. Second costal section variable but usually shorter. Mid tibia with at least 1 long posterodorsal seta, i.e. mid tibia with long paired setae. Male mid tibia with or without long ventroapical spur.

39 (42) Ventroapical seta (spur) of mid tibia very long, ca. half as long as mid basitarsus. Tibiae without light rings, at most apices lighter.

40 (41) Male mid tibia ventrally without long hairs. Surstylus long with an additional black lateral thorn on caudal lobe ( DEEMING 1969: fig. 19). India P. brunettii ( DEEMING, 1969)

41 (40) Male mid tibia with extremely long upright hairs. Surstylus ( Fig. 25 )

shorter with several thick setae on caudal lobe. Taiwan

P. longicalcar sp. n., pro parte

Aberrant males without an appendage on R 2+3 run here.

42 (39) Mid tibia with or without long ventroapical spur; if present, much shorter than half length of mid basitarsus.

43 (50) Male tergite 5 broadened on right side with 9–13 extremely long thick marginal bristles. Posterior lobe of surstylus without apical tooth ( Fig. 49 ). Vein R 4+5 slightly bent towards costa.

44 (45) Male fore tibia without long hairs anteroventrally (hairs shorter than tibial diameter). Male mid tibia with medium-long hairs ventrally. Sri Lanka and Malaysia P. pappi HAYASHI, 1997

45 (44) Male fore tibia with extremely long hairs (some of the hairs longer than 1/3 length of fore tibia).

46 (47) Male fore tibia with extremely long hairs restricted to basal third of tibia. Male mid tibia ventrally with short (but thicker, stiff) hairs only. Biak I. and Papua New Guinea P. insularis HAYASHI, 1997

47 (46) Male fore tibia with extremely long hairs along its entire length ( Fig. 48 ). Male mid tibia ventrally with long hairs (longer than tibial diameter).

48 (49) Hairs on male fore tibia at most slightly longer than 1/3 length of tibia (see fig. 5 of HAYASHI 1997). Anterior lobe of surstylus with shorter hairs ( HAYASHI 1997: fig. 11), posterior lobe comparatively smaller. Christmas Island P. pectiniterga ( DEEMING, 1964)

49 (48) Longest hairs on male fore tibia as long as 2/3 length of tibia ( Fig. 48 ). Anterior lobe of surstylus with very long hairs ( Fig. 49 ), posterior lobe larger than in P. pectiniterga . Philippines P. pilipino sp. n.

50 (43) Male tergite 5 normal, with the usual marginal bristles.

51 (52) Male mid tibia apically curved inwards and dorsoventrally flattened, ventrally with medium-long hairs. Male mid tibia apically without va seta but with a patch of short black spinules. Male fore tibia ventrally and posteriorly, as well as all fore tarsomeres with extremely long hairs. Large species, wing length 3.15 mm. Taiwan P. curvipes sp. n.

52 (51) Male mid tibia not curved inwards and not flattened. Usually smaller species.

53 (56) Apical half of vein R 4+5 straight, medial part more strongly curved.

54 (55) Male mid tibia ventrally with very long hairs in distal 1/3 ( DEEMING 1969: fig. 4); some of the ventral hairs thicker than adjacent ones. Male fore tibia and basitarsus posteriorly and posteroventrally with hairs shorter than diameter of basitarsus. Posterior lobe (ventromedial process) of male surstylus with a very long droplet-shaped apical tooth ( DEEMING 1969: fig. 16). Nepal P. himalayensis ( DEEMING, 1969)

55 (54) Male mid tibia ventrally with shorter hairs (only some posterior hairs longer than tibial diameter). Male fore tibia and basitarsus with hairs much longer than diameter of tibia/basitarsus at their bases. Posterior lobe (ventromedial process) of male surstylus ( Fig. 67 ) with an extremely long, strap-like black spine. Sumatra P. subpilimana sp. n.

56 (53) Vein R 4+5 more strongly bent towards the costa, i.e. apical part not straight, consequently medial part less curved (basal half slightly S-shaped).

57 (66) Male mid tibia ventrally with long hairs, no distinct, strong ventroapical seta but ventroapically at most with a transverse row of more or less curved, longer hairs instead. Male fore tibia ventrally and anteriorly, as well as fore basitarsus, or even other tarsomeres, with long hairs.

58 (59) Subepandrial sclerite ventrally with 6–7 long thick thorns ( Fig. 51 ). Surstylar lobes very short, surstylus basally (caudally) with numerous short thick setae ( Fig. 52 ). Sabah P. sabahi sp. n.

59 (58) Subepandrial sclerite normal, i.e. without long thick thorns ventrally. Surstylus otherwise shaped.

60 (63) Male mid tibia with ventral hairs strong, straight (stiff) but not longer than tibial diameter (at most some longer hairs subapically), some of the hairs blunt and thicker than adjacent hairs. Usually 4 short posterodorsals plus a longer preapical posterodorsal seta.

61 (62) Male ventral sclerites posterior to sternite 5 form two pairs of hairy processes ( PAPP 1991 a: fig. 26). Male surstylus ( PAPP 1991 a: fig. 30). Widespread P. punctipennis ( WIEDEMANN, 1824)

62 (61) Male postabdomen with a single medial T or wide Y-shaped process ( HAYASHI 2002). Male surstylus ( HAYASHI 2002). Widespread P. affinis HAYASHI, 2002

63 (60) Male mid tibia with thinner but longer hairs, numerous hairs longer than tibial diameter; dorsal armature may be different.

64 (65) Male mid tibia ventroapically with a transverse row of 4–5 curved black thornlets. Surstylus without a large lateral process. Taiwan, Sumatra P. amputata ( DUDA, 1925)

65 (64) Ventroapical part of male mid tibia with a patch of ca. 10 short black setulae. Epandrium dorsally with a pair of very long bristles. Surstylus ( Fig. 43 ) laterally with an extremely long curved, posteriorly directed spine. Sumatra P. pilimana sp. n.

66 (57) Male mid tibia ventrally without long hairs, ventroapical seta more or less strong or mid basitarsus modified.

67 (72) Male fore tibia and basitarsus without long hairs. Female mid tibia with two short ventroapicals, which are much shorter than tibial diameter subapically.

68 (69) Male fore tibia ventrally with a row of black spines ( Fig. 58 ). Surstylus ( Figs 61–62, 64 ) with a pecten of 11 teeth. Sumatra P. spinipes sp. n.

69 (68) Male fore tibia ventrally without black spines.

70 (71) Male and female basitarsi normal. Vein R 2+3 apically angularly bent to costa. Costa terminates at tipof vein R 4+5. Sumatra, Java, Taiwan,? Burma P. rectinervis ( DUDA, 1925)

71 (70) Male mid basitarsus with dense black setulae on its whole ventral surface.

Female fore basitarsus, as well as hind 2nd tarsomere with short but very dense hairs; setulae on ventral surface of mid basitarsus also denser than usual. Vein R 2+3 bent along a wide arc to costa. Costa overruns tipof vein

R 4+5 by a length equalling three times its diameter. Sumatra

P. peniculifera sp. n.

72 (67) Male fore tibia and basitarsus (possibly other tarsomeres) posteriorly, or ventrally to posteriorly, with long hairs (longer than their diameter at bases of hairs).

73 (74) No yellow ring on middle of fore tibia. Facial plate not much protruding. Face, frons and genae dark brown. Hairs on male fore tibia and tarsomeres very long (some hairs more than twice longer than tibial diameter). Taiwan P. nigrotibiata ( DUDA, 1925)

74 (73) A yellow ring on middle of fore tibia. Facial plate more convex. At least face or frons reddish or even lighter.

75 (76) Subepandrial sclerite laterally with (3-)4 long thick thorns ( Figs 56–57 ). Surstylus ( Fig. 54 ) medially with a thick but acute thorn and several strong setae. Postgonite ( Fig. 55 ) strongly curved and rather broad. Apical part of first flagellomere markedly lighter than rest of flagellomere, also in females. Philippines, Tonga and Society Is. P. spinicauda sp. n.

76 (75) Subepandrial sclerite without thick thorns. Surstylus different.

77 (80) Male subepandrial sclerite with a pair of peculiar processes ( Fig. 32 ). Surstylus trilobed with several thick thorns.

78 (79) Male subepandrial sclerite with a pair of large, broad, upcurved processes ( PAPP 1991 a: figs 33, 35). Postgonite ( PAPP 1991 a: fig. 34) geniculately bent, apical part not very broad. Widespread P. varians ( DUDA, 1925)

79 (78) Male subepandrial sclerite with a pair of acutely pointed straight processes ( Fig. 32 ). Postgonite ( Fig. 31 ) bisinuate with broader, partly membraneous, apical part. Sri Lanka, Taiwan, probably widespread

P. nigra sp. n.

80 (77) Male subepandrial sclerite simple ( Fig. 39 ). Surstylus ( Fig. 41 ) bilobed.

81 (82) Medial part of R 2+3 slightly concave, i.e. no section parallel with costa. Posteromedial lobe of surstylus shorter and broader with a laterally placed thorn ( PAPP 1991 a: fig. 13). Postgonite ( PAPP 1991 a: fig. 15) less broad, caudal margin angulate. Subepandrial sclerite ( PAPP 1991 a: fig. 11). Taiwan P. cryptica L. PAPP, 1991

82 (81) A rather long medial part of R 2+3 parallel to costa, apical part bent along a wide arc (i.e. not angulately) into costa. Posteromedial lobe of surstylus longer, apically with a bidentate thorn ( Fig. 41 ). Postgonite ( Fig. 40 ) very broad, caudal margin arcuately bent. Subepandrial sclerite ( Fig. 39 ). Sabah P. paracryptica sp. n.

Acknowledgements – I am grateful to Dr ALLEN L. NORRBOM ( SEL, USDA, c/o National Museum of Natural History / Smithsonian Institution , Washington), Dr CHEN W. YOUNG ( The Carnegie Museum of Natural History , Section of Invertebrate Zoology , Pittsburgh, PA, USA) and JOHN SWANN ( Royal Ontario Museum ) for the loans of the Poecilosomella specimens. It is with pleasure that I acknowledge Dr HERMAN DE JONG ( Zoölogisch Museum , University of Amsterdam ) and Dr NIGEL WYATT (The Natural History Museum London) for information on type specimens .

This paper belongs to a series resulting from the joint project of the Hungarian Natural History Museum and the Taiwan Forestry Research Institute, entitled “Biodiversity studies on the insect fauna of Taiwan: taxonomy, faunagenetics and inventory of selected insect families”, No. 5.

Figures

Figs 65–68. Poecilosomella subpilimana sp. n., paratype male, postabdomen and genitalia. 65 = sternites 5 and6, ventral view,66 =subepandrial sclerite, caudal view, 67 = surstylus, lateral view,68

Figs 10–15. Poecilosomella spp.males.10–13 =P.ornata (DE MEIJERE), Taiwan: 10 = sternite 5 and 6, ventral view, 11 = left half of subepandrial sclerite, caudal view; 12 = surstylus, lateral view, 13 = postgonite, lateral view. 14–15 = P. meijerei (DUDA): 14 = surstylus, true lateral view, 15 =

Figs 69–72. Poecilosomella picturata (MALLOCH), holotype male. 69 = right half of subepandrial sclerite, caudal view, 70 = surstylus, true lateral view, 71 = postgonite, lateral view, 72 = sternites 5

Figs 20–23. Poecilosomella formosana sp. n., paratype male. 20 = right half of subepandrial sclerite, caudal view, 21 = surstylus, true lateral view, 22 = sternite 5 and 6, ventral view, 23 = postgonite, lat-

Figs 1–5. Poecilosomella borboroides (WALKER), male postabdomen and genitalia. 1 = sternite 5 and 6, ventral view, 2 = right half of subepandrial sclerite, caudal view, 3 = postgonite, 4 = surstylus, lateral view (Sabah), 5 = apical part of surstylus (India). Scales: 0.2 mm for Fig. 1, 0.1 mm for Figs

Figs 6–9.Poecilosomella borborus sp. n., paratype male, postabdomen and genitalia. 6 = right half of subepandrial sclerite, caudal view, 7 = surstylus, lateral view, 8 = postgonite in widest extension, i.e. not lateral view, 9 = sternite 5 and pregenital sternites, ventral view. Scales: 0.2 mm for Fig. 9, 0.1

Figs 24–27. Poecilosomella longicalcar sp. n., paratype male, terminalia. 24 = subepandrial sclerite, caudal view, 25 = surstylus, lateral view, 26 = sternites 5 and6, ventral view, 27 = postgonite, true lat-

Figs 46–49. Poecilosomella pilipino sp. n., paratype male. 46 = genital complex with postgonite, lateral view, 47 = postgonite in higher magnification, lateral view, 48 = fore tibia, anterior view, 49 =

Figs 50–53. Poecilosomella sabahi sp. n., paratype male, postabdomen and genitalia.50 = sternites 5 and 6, ventral view (S5 setae omitted), 51 = right half of subepandrial sclerite, caudal view, 52 = surstylus, lateralview,53 =postgonite,lateralview.Scales:0.2mm for Figs50–52, 0.1mm for Fig.53

Figs 42–45. Poecilosomella pilimana sp. n., paratype male, postabdomen and genitalia. 42 = subepandrial sclerite, caudal view, 43 = surstylus, lateral view, 44 = sternites 5 and 6, subventral

Figs 58–64. Poecilosomella spinipes sp. n., paratype male. 58 = fore tibia with basitarsus, lateral view, 59 = basal abdominal tergites, dorsal view, 60 = sternite 5 (medially torn asunder) with pregenital sternites (moved with the right half of S5), ventral view, 61 = surstylus, lateral (outer) view, 62 = same, inner view, 63 = postgonite, lateral view, 64 = apex of surstylus in dorsal view of pecten. Scales:0.4mm for Fig.59,0.2 mm forFig.60, 0.1 mm forFigs 61–64 (for Fig.58 same asfor Fig.48)

Figs 54–57. Poecilosomella spinicauda sp. n., paratype male, genitalia. 54 = surstylus, outer widest (subcaudal) view, 55 = postgonite, lateral view, 56 = ventral part of epandrium, lateral view (as if thorns on subepandrial sclerite were on epandrium), 57 = left half of subepandrial sclerite, subventral

Figs 31–34. Poecilosomella nigra sp.n., paratype male.31 = genital complex with postgonite, lateral view, 32 = right half of subepandrial sclerite with apex of epandrium, 33 = surstylus, outer lateral

Figs 39–41. Poecilosomella paracryptica sp. n., paratype male, postabdomen and genitalia. 39 = subepandrial sclerite, caudal view, 40 = postgonite, lateral view, 41 = surstylus, lateral view. Scale: