Phonotimpus boneti, Platnick & Chamé-Vázquez & Ibarra-Núñez, 2022

Platnick, Norman I., Chamé-Vázquez, David & Ibarra-Núñez, Guillermo, 2022, The guardstone spiders of the genus Phonotimpus Gertsch & Davis (Araneae: Phrurolithidae) from northeastern Mexico, Zootaxa 5219 (1), pp. 1-48 : 19-20

Phonotimpus boneti sp. nov.

Figures 126–133 , 156–162

Type material. Female holotype from Chipinque, Monterrey, Nuevo León, Mexico (Sept. 25, 1943; F. Bonet), deposited in AMNH .

Other material examined. MEXICO: Nuevo León: Chipinque Mesa, Monterrey , elev. 1646 m, June 21–25, 1969, human dung trap, forest, S., J. Peck, 1♂ ( AMNH) .

Etymology. The specific name is a patronym in honor of the collector of the holotype and many other significant specimens of the genus.

Diagnosis. The female resembles P. valles sp. nov. by having copulatory openings on the anterior border of the atrium, copulatory ducts laterally directed, and tubular primary spermathecae, but the atrium has a shallow, median septum, and a distinctive anterior, transverse epigynal hood, with posterior rim sclerotized ( Figs 160–162 ). The only available male is in poor condition but can be readily separated by having the retrolateral tibial apophysis longer and much larger than the dorsal apophysis, tip hooked ( Figs 157–159 ), and apical tegular apophysis pointing to distalmargin ( Fig. 157 ). The male was not collected with the female, but both were collected in the same locality.

Description. Male (Chipinque): Total length 1.68. Specimen darkened, poorly preserved, original coloration uncertain, pars cephalica sloping evenly from elevated pars thoracica to clypeus, each cheliceral paturon with one erect bristle near base; abdomen narrow, coloration uniformly faded, dorsal scutum shiny, rectangular, occupying about two thirds of abdominal length, half width ( Figs 126–129 ). Legs darkened from poor preservation, markings indeterminate. Leg spination: femora: I d1-0-0, p0-0-2; II–IV d1-0-0; tibiae: I v4-4-4; II v4-2-2; metatarsi I, II v2-2-2.

Palpal femur club-shaped, with one dorso-distal spine, retrolateral excavation shallow but extending over twothirds of femur length, patch of spinules on distal, prolateral half; dorsal tibial apophysis distally attenuate, shorter and thinner than retrolateral tibial apophysis, which is long, massive, base occupying full length of tibia at that point, inner margin sinuous, with subdistal expansion, tip strongly hooked ( Figs 157–159 ); distal border of palpal tibia deeply notched prolaterally ( Figs 156–157 ); very shallow groove on middle of cymbium, corresponding with dorsal tibial apophysis ( Fig. 159 ); tegulum expanded proximally; base of embolus slender, membranous, occupying about one-fifth of bulb width; embolus short, tubular, weak, distally directed; embolar basal process slender; conductor narrow, tip slightly widened; apical tegular apophysis rod-shaped, tip blunt, pointing to distal-retrolateral margin ( Fig. 157 ).

Female (holotype): Total length 1.80. Carapace yellow, with undarkened lateral margins, dusky markings restricted to U-shaped wide band situated about halfway between lateral margins and unmarked median area, pars thoracica steeply sloping, pars cephalica as in male; sternum, mouthparts yellow, unmarked, each cheliceral paturon slightly darkened, with two erect bristles near base; abdomen narrow, dorsum light gray, with indistinct cardiac markings, posteriorly with five white chevrons, most anterior and most posterior ones largest, extending fully to sides, weak, shiny scutum triangular, covering only cardiac area, sides, venter pale yellow, unmarked except for dark circle around spinnerets ( Figs 130–133 ); legs yellow, all femora, patellae, tibiae, metatarsi each with distal dark markings. Leg spination as in male except tibia I v4-4-4; II v4-2-2-1p; metatarsi I, II v2-2-2- 1p.

Epigynum large, transverse hood with sclerotized posterior rim, overhanging a deep, rounded atrium, with low, longitudinal septum; small copulatory openings situated on each side of septum; copulatory ducts short, sclerotized, laterally oriented; bursae globose, almost contiguous distally; primary spermathecae tube-shaped, thickened distally, about as long as bursae, posteriorly directed, extending almost to posterior epigynal margin; secondary spermathecae tiny, hook-shaped ( Figs 160–162 ).

Distribution. Known only from the Municipio de Monterrey in northern Nuevo León, where it is sympatric with P. chipinque sp. nov. ( Fig. 339 ).

Figures

FIGURE 339. Northeast Mexico, showing distribution of the Phonotimpus species described in this work.

FIGURES 122–141. 122–125. Phonotimpus arcitos sp. nov., female. 126–133. P. boneti sp. nov., male (126–129) and female (130–133). 134–141. P. chipinque sp. nov., male (134–137) and female (138–141). 122, 126, 130, 134, 138. Carapace, dorsal view. 124, 128, 132, 136, 140. Same, ventral view. 123, 127, 131, 135, 139. Abdomen, dorsal view. 125, 129, 133, 137, 141. Same, ventral view.

FIGURES 156–162. Phonotimpus boneti sp. nov., male (156–159) and female (160–162). 156. Left palp, prolateral view. 157. Same, ventral view. 158. Same, retrolateral view. 159. Same, dorsal view. 160. Epigynum, ventral view. 161. Cleared epigynum, ventral view. 162. Same, dorsal view.