Chelipoda, MACQUART, 1823

Plant, Adrian R., 2009, Diversity Of Chelipoda Macquart, 1823 (Diptera: Empididae: Hemerodromiinae) In Northern Thailand With Discussion Of A Biodiversity ‘ Hot Spot’ At Doi Inthanon, Raffles Bulletin of Zoology 57 (2), pp. 255-277 : 256

Key to species of Chelipoda from Northern Thailand

1. Wing with strong black subterminal band commencing at apex of cell dm leaving apex of wing clear .................................... ........................... C. menglunana Grootaert, Yang & Saigusa

– Wing weakly pigmented; if darkened, never with distinct subterminal band .................................................................. 2

2 Legs conspicuously slender; F 1 8–9x long as wide ( Fig. 26 ) with a single row of denticles between double rows of ventral spines ........................................ C. macrosceles , new species

– Legs stouter ( Figs. 25, 27 ); F 1 at most 6x long as wide, usually strongly inflated (F 1 3.4 –4.8x long as wide), a double row of denticles between double rows of ventral spines (if only single row of denticles present then F 1 conspicuously inflated) .. 3

3 Ground colour of thorax yellow or brownish yellow at least on pleura; scutum sometimes darkened or with darker stripes ............................................................................................... 4

– Ground colour of thorax black, at most with propleuron contrastingly yellowish ...................................................... 12

4 Head yellow ......................................................................... 5

– Head black ............................................................................ 6

5 Upper occiput, vertex and frons darkened; scutum yellow with dark median stripe ( Fig. 21 ) .................... C. flavida Brunetti

– Upper occiput, vertex and frons yellow; scutum yellow without dark median stripe .................... C. manggawna , new species

6 Thoracic dorsum brownish or yellowish black ................... 7

– Thoracic dorsum yellow, sometimes with dark median stripe ............................................................................................... 9

7 Front coxa without distinct anterobasal spine. Antenna with basal segments dark reddish yellow, not contrasting strongly with dark postpedicel ........... C. guangxiensis Yang & Yang

– Front coxa with distinct anterobasal spine. Antenna with basal segments yellowish, contrasting with dark postpedicel ...... 8

8 Scutum diffusely edged paler brown laterally, not distinctly contrasting with darker brown central area ( Fig. 23 ) ............. ................... C. nakropa , new species (mid elevation morph)

– Scutum sharply edged yellowish lateral to line of dorsocentrals, the yellow margins contrasting with central broad, almost blackish area ( Fig. 22 ) ............................................................. ................... C. nakropa , new species (high elevation morph)

9 Scutum more or less wholly yellow .................................. 10

– Scutum yellow with dark median stripe ........................... 11

10 Front coxa with distinct anterobasal spine. F 1 with single row of denticles. Front tarsomeres 2 and 3 distinctly spinose ...... .................................................... C. kameawuta , new species

– Front coxa without distinct anteroventral spine. F 1 with double row of denticles. Front tarsomeres 2 and 3 not distinctly spinose ................................... C. thaosuranaria , new species

11 Scutum with narrow dark median stripe ( Fig. 24 ). A contrasting deep black mark between and behind front coxae on av margin of katepisternum ........................ C. chaiamnata , new species

– Scutum with broader dark median stripe. Area between and behind front coxae yellow, not contrasting with surroundings ................................................................ Chelipoda species D

12 Propleuron blackish (at most faintly yellowish), not contrasting with rest of pleura .............................................................. 13

– Propleuron yellowish or reddish, contrasting with rest of black pleura .................................................................................. 14

13 Thoracic setae and spines on F 1 dark. Male sternites 7 & 8 with sparse black setae ............ C. hubeiensis Yang & Yang

– Thoracic setae and spines on F 1 yellow. Male sternites 7 & 8 with numerous yellow setae ( Fig. 11 ) ..................................... ................................................ C. meenamluang , new species

14 Front coxa without distinct anterobasal spine. Posterior dc (in line with npl) about as strong as anterior dc .......................... ....................................................... C. nakladam , new species

– Front coxa with distinct anterobasal spine. Posterior dc (in line with npl) minute, much smaller than anterior dc ............. 15

15 Male cercus with strong pointed ad process ( Fig. 4 ). Female basal antennal segments yellowish ..... C. inthawichayanona , new species

– Male cercus with strong pointed ad process and lower less pointed lobe ( Fig. 8 ) ........................ C. laisoma , new species

Figures

Figs. 25–28. Chelipoda species: 25. C. manggawna new species, male; 26. C. macrosceles new species, female habitus; 27. C. nakropa new species, male, front femora showing chaetotaxy; 28. C. hubeiensis Yang & Yang, female, wing. Abbreviations: bm, cell bm; br, cell br; bs, basal spines; d, denticles; dm, cell dm; dm-cu, crossvein dm-cu; s, spines.

Figs. 19–24. Thoracic dorsum of Chelipoda spp. showing schematic pattern of ground colour. 19–20. C. macrosceles new species: 19. male; 20. female: 21. C. flavida Brunneti, male: 22–23. C. nakropa new species, male: 22. high elevation morph; 23. mid elevation morph: 24. C. chaiamnata new species, male.

Figs. 7–11. Male genitalia of Chelipoda species in lateral view: 7. C. kameawuta new species; 8. C. laisoma new species; 9. C. macrosceles new species; 10. C. manggawna new species; 11. C. meenamluang new species.

Figs. 1–6. Male genitalia of Chelipoda species in lateral view: 1. C. chaiamnata new species; 2. C. flavida Brunetti; 3. C. guangxiensis Yang & Yang; 4. C. inthawichayanona new species: 5–6 C. hubeiensis Yang & Yang; 5. typical form; 6. variant (outline only).