Exocelina bacchusi (Balke, 1998)

Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael, 2019, Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 878, pp. 73-143 : 73

6. Exocelina bacchusi (Balke, 1998) Figs 16 , 22 , 24

Copelatus (Papuadytes) bacchusi Balke, 1998: 326; Nilsson 2001: 76 (catalogue).

Papuadytes bacchusi (Balke, 1998): Nilsson and Fery 2006: 56 (comb. nov.).

Exocelina bacchusi (Balke, 1998): Nilsson 2007: 33 (comb. nov.).

Exocelina bacchusi MB1521: Toussaint et al. 2015: supplementary figs S1, S2, tab. S3, and information S5, S6.

Exocelina undescribed sp. MB0257: Toussaint et al. 2014: supplementary figs 1-4, tab. 2; Toussaint et al. 2015: supplementary figs S1, S2, tab. S3, and information S5, S6.

Type locality.

Papua New Guinea: Madang Province, Finisterre Range, Damanti, 05°53'26.5"S, 145°57'50.6"E, 1180 m a.s.l.

Type material studied.

Holotype: male "Stn. No. 39", "NEW GUINEA: Madang Dist., Finisterre Mts. Damanti 3,550 ft. 2 –11.x.1964.”, "M.E. Bacchus. B.M. 1965-120", “HOLOTYPUS” [red], " Copelatus bacchusi Balke des. 1997" [red] (BMNH). Paratypes: 1 male with the same label as the holotype and additionally with red label "Paratypus Copelatus bacchusi Balke des. 1997" (BMNH). 2 males "Stn. No. 49", "NEW GUINEA: Madang Dist., Finisterre Mts. Budemu c. 4000 ft. 15 –24.x.1964.”, "M.E. Bacchus. B.M. 1965-120" (BMNH, NHMW). 4 females "Stn. No. 74", "NEW GUINEA: Madang Dist., Finisterre Mts. Budemu c. 4000 ft. 15 –24.x.1964.”, "M.E. Bacchus. B.M. 1965-120" (BMNH, NHMW). Note: in the original description (Balke, 1998: 326), the paratypes with the same label as the holotype and from locality "Stn. No. 49" were erroneously indicated as females.

Additional material.

Madang: 2 males "Papua New Guinea: Madang, Simbai area, 1200m, 11.iii.2007, 05.13.333S 144.37.611E, Kinibel (PNG 153) (ZSM). 15 males, 6 females "Papua New Guinea: Madang, Simbai-Mombeen, 1100m, 11.iii.2007, 05.12.876S 144.41.759E, Kinibel (PNG 154), one male with an additional green label "DNA M.Balke 3318" (NHMW, ZSM). 1 male, 14 females "Papua New Guinea: Madang, Keki-Sewan, Adalbert [sic!] Mts., 700m, 30.xi.2006, nr 04.41.802S 145.25.460E, Binatang Boys (PNG 120)" (ZSM). 4 males, 4 females "Papua New Guinea: Madang, Adalbert [sic!] Mts., creek nr Keki, 790m, 28.xi.1994, 04.42.300S 145.25.089E, Binatang Boys leg. (PNG 53a)" (NHMW, ZSM). 2 males, 5 females "Papua New Guinea: Madang, Adalbert [sic!] Mts., Keki, 850m, 4.v.2006, nr 04.42.300S 145.25.089E, Manaono leg. (PNG 52)" (ZSM). 83 males, 84 females "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 1200m, 24.xi.2006, nr 05.52.754S 145.23.209E, Balke & Kinibel (PNG 110)", one of them with an additional green label "DNA M.Balke 1521" (NHMW, ZSM). 7 males, 3 femlaes "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 1500m, 24.xi.2006, 05.51.964S 145.23.604E, Balke & Kinibel (PNG 111)" (ZSM). 3 females "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 800m, 24.xi.2006, 05.50.021S 145.24.664E, Balke & Kinibel (PNG 112)" (ZSM). 1 male, 1 female "Papua New Guinea: Madang, Akameku - Brahmin, Bismarck Range, 750m, 25.xi.2006, 05.49.892S 145.24.491E, Balke & Kinibel (PNG 113)" (ZSM). 1 male "Ibisca Niugini, PNG 26-28.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-P-1/8-d02 / Plot 16 / P1608 Vial 18767" (ZSM). 1 male "Ibisca Niugini, PNG 26-28.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-T-1/8-d02 / Plot 20 / P1640 Vial 18781" (ZSM). 3 females "Ibisca Niugini, PNG 26-28.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-P-1/8-d02 / Plot 16 / P1608 Vial 18767" (ZSM). 1 female "Ibisca Niugini, PNG 26-28.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-O-1/8-d02 / Plot 15 / P1600 Vial 18763" (ZSM). 1 female "Ibisca Niugini, PNG 26-28.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-K-1/8-d02 / Plot 11 / P1568 Vial 17122" (ZSM). 1 female "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-A-2/8-d03 / Plot 1 / P1489 Vial 17237" (ZSM). 1 female "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-C-2/8-d03 / Plot 3 / P1505 Vial 17179" (ZSM). 6 females "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1529 Vial 16853" (ZSM). 1 male, 4 females "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m -5,720873833 145,269470 2 MW1200 / P1553 Vial 09007" (ZSM). 1 male, 4 females "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1545 Vial 16863" (ZSM). 1 male, 7 females "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1561 Vial 16873" (ZSM). 1 male "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-A-2/8-d03 / Plot 1 / P1489 Vial 17237" (ZSM). 7 females "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-E-2/8-d03 / Plot 5 / P1521 Vial 17210" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1569 Vial 17302" (ZSM). 3 females "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1601 Vial 17313" (ZSM). 11 females "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1595 Vial 18799" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1593 Vial 17462" (ZSM). 2 males "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1577 Vial 18802" (ZSM). 1 male "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1625 Vial 18821" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1633 Vial 18848" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1617 Vial 18813" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1601 Vial 17313" (ZSM). 2 females "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1585 Vial 18825" (ZSM). 1 male, 13 females "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1609 Vial 18855" (ZSM). 1 female "Ibisca Niugini, PNG 29-31.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-D-3/8-d05 / Plot 4 / P1514 Vial 16946" (ZSM). 1 female "Ibisca Niugini, PNG 29-31.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-F-3/8-d05 / Plot 6 / P1530 Vial 16936" (ZSM). 1 female "Ibisca Niugini, PNG 29-31.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-A-3/8-d05 / Plot 1 / P1490 Vial 16931" (ZSM). 2 females "Ibisca Niugini, PNG 29-31.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-H-3/8-d05 / Plot 8 / P1546 Vial 17249" (ZSM). 1 female "Ibisca Niugini, PNG 29-31.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-C-3/8-d05 / Plot 3 / P1506 Vial 17235" (ZSM). 1 female "Ibisca Niugini, PNG 29-31.x.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-J-3/8-d05 / Plot 10 / P1562 Vial 16881" (ZSM). 2 females "Ibisca Niugini, PNG 30.x.-1.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1578 Vial 17625" (ZSM). 4 females "Ibisca Niugini, PNG 30.x.-1.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1570 Vial 17585" (ZSM). 1 female "Ibisca Niugini, PNG 30.x.-1.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1634 Vial 17605" (ZSM). 1 female "Ibisca Niugini, PNG 31.x.-2.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1547 Vial 17303" (ZSM). 1 female "Ibisca Niugini, PNG 31.x.-2.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1515 Vial 17326" (ZSM). 1 female "Ibisca Niugini, PNG 31.x.-2.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1491 Vial 17331" (ZSM). 1 female "Ibisca Niugini, PNG 31.x.-2.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1531 Vial 17347" (ZSM). 1 female "Ibisca Niugini, PNG 31.x.-2.xi.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-E-4/8-d07 / Plot 5 / P1523 Vial 17348" (ZSM). 2 females "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1579 Vial 18787" (ZSM). 8 females "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1643 Vial 18794" (ZSM). 1 female "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1595 Vial 18799" (ZSM). 3 females "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1571 Vial 16947" (ZSM). 1 female "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1635 Vial 16968" (ZSM). 1 female "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-S-4/8-d08 / Plot 19 / P1635 Vial 16968-CODYTI" (ZSM). 1 male, 10 females "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1611 Vial 16950" (ZSM). 2 females "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1611 Vial 16950" (ZSM). 2 females "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1635 Vial 16968" (ZSM). 1 female "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1516 Vial 17287" (ZSM). 2 females "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1548 Vial 17297" (ZSM). 2 females "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1532 Vial 17314" (ZSM). 1 female "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1492 Vial 17355" (ZSM). 1 male, 2 females "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1564 Vial 16857" (ZSM). 1 female "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m", " -5,72090292 145,2714691 FIT-MW1200-C-5/8-d09 / Plot 3 / P1508 Vial 14052-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m", " -5,72090292 145,2714691 FIT-MW1200-I-5/8-d09 / Plot 9 / P1556 Vial 17374" (ZSM). 1 female "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 ", "FIT-MW1200-E-5/8-d09 / Plot 5 / P1524 Vial 16861-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 3-5.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1636 Vial 17324" (ZSM). 2 males "Ibisca Niugini, PNG 3-5.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1612 Vial 17292" (ZSM). 1 male "Ibisca Niugini, PNG 3-5.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1572 Vial 18837" (ZSM). 1 female "Ibisca Niugini, PNG 3-5.xi.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-O-5/8-d10 / Plot 15 / P1604 Vial 17325" (ZSM). 1 male "Ibisca Niugini, PNG 4-6.xi.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-F-6/8-d11 / Plot 6 / P1533 Vial 17257" (ZSM). 1 male "Ibisca Niugini, PNG 5-7.xi.2012 Mount Wilhelm 1200m", " -5,720873833 145,2694702 FIT-MW1200-K-6/8-d12 / Plot 11 / P1573 Vial 17082" (ZSM). 1 female "Ibisca Niugini, PNG 8-10.xi.2012 Mount Wilhelm 1200m -5,721022129 145,2703094 MW1200 / P1503 Vial 16995" (ZSM). 1 female "Ibisca Niugini, PNG 8-10.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1527 Vial 16879" (ZSM). 1 female "Ibisca Niugini, PNG 8-10.xi.2012 Mount Wilhelm 1200m -5,720873833 145,2694702 MW1200 / P1503 Vial 16995" (ZSM). 1 male, 1 female "Ibisca Niugini, PNG 9-11.xi.2012 Mount Wilhelm 1200m," " -5,720873833 145,2694702 FIT-MW1200-T-8/8-d16 / Plot 20 / P1647 Vial 17039-CODYTI" (ZSM). 4 females "Ibisca Niugini, PNG 9-11.xi.2012 Mount Wilhelm 1200m," " -5,720873833 145,2694702 FIT-MW1200-P-8/8-d16 / Plot 16 / P1615 Vial 17012-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 9-11.xi.2012 Mount Wilhelm 1200m," " -5,720873833 145,2694702 FIT-MW1200-M-8/8-d16 / Plot 13 / P1591 Vial 17035-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 1-3.xi.2012 Mount Wilhelm 1700m -5,79269238 145,235611 MW1700 / P1961 Vial 06629" (ZSM). Eastern Highlands: 3 male, 3 females "Papua New Guinea: Eastern Highlands, Bena Bridge, 1400m, 8.xii.2007, 06.10.781S 145.26.034E, Balke & Sagata (PNG 164)" (ZSM). Simbu/Eastern Highlands: 2 males "Papua New Guinea: Simbu/EHPr. Crater Mountain, Sera - Herowana, Wara Pima, 900 m, 15IX2002, Balke & Sagata (PNG 011)" (ZSM). 16 males "Papua New Guinea: Crater Mountain, Sera - Herowana, upper Oh River, 1200 m, 15IX2002, Balke & Sagata (PNG 012)" (NHMW, ZSM). 2 males, 4 females "Papua New Guinea: Crater Mountain, Sera - Herowana, Jau river, 1100m, 15IX2002, Balke & Sagata (PNG 013)" (ZSM). 12 males "Papua New Guinea: Simbu/EHPr. Crater Mountain, Sera - Herowana, Jau river, 1000 m, 15IX2002, Balke & Sagata (PNG 015)" (NHMW, ZSM). 1 male, 3 females "Papua New Guinea: Simbu / EHP, Crater Mountain, Sera - Herowana, Sima river, 1250m, 15IX2002, Balke & Sagata (PNG 016)" (ZSM). Simbu: 5 males "Papua New Guinea: Supa Haia, 1023m, 10.ix.2002, K.Sagata (WB1)" (NHMW, ZSM). 1 male "Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 600m, 12IX2002, Balke & Sagata, (PNG 003)" (ZSM). 3 males "Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 500m, 12IX2002, Balke & Sagata, (PNG 005)" (ZSM). 1 male "Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 500m, 12IX2002, Balke & Sagata, (PNG 006)" (ZSM). 11 males, 14 females "Papua New Guinea: Simbu/EHPr. Crater Mountain, Wara Sera Station, 800 m, 14IX2002, Balke & Sagata (PNG 009)" (NHMW, ZSM). 12 males, 7 females "Papua New Guinea: Simbu/EHPr. Crater Mountain, Wara Sera Station, 800 m, 14IX2002, Balke & Sagata (PNG 10)", one male with additional labels "257 DNA M Balke" [green], "sp.17 SEM 19" (ZSM). 27 males, 31 females "Papua New Guinea: Crater Mountain, Wara Sera Station, 800 m, 14IX2002, Balke & Sagata (PNG 010)" (NHMW, ZSM). Morobe: 62 males, 27 females "PAPUA N.G.: Morobe Prov. E Pindiu, Kobau 24.4.1998, 1400 m, leg. A. Riedel" (NHMW, ZSM). 1 male "Papua New Guinea: Morobe, Pindiu, Sulemana, 850 m, 15.x.2009, 06.25.169S 147.32.11E, Inaho (08) (PNG 208)", "DNA M.Balke 3825" [green] (ZSM). 2 males "PNG: Huon Peninsula, Morobe Prov., Yus conservation area [5°53'54"S, 146°48'15"E], 1398m, 24.viii.2010, Bega", "DNA M. Balke 6531" [green text], "DNA M. Balke 6532" [green text] (ZSM). 6 females "PNG: Huon Peninsula, Morobe Prov., Yus conservation area (Y7), 1398m, 24.viii.2010, Huon, Bega" (ZSM). Gulf: 9 males, 8 females "Papua New Guinea: Gulf, Marawaka, nr Ande, 1000m, 10.xi.2006, 07.03.598S 145.44.375E, Balke & Kinibel (PNG 89)" (NHMW, ZSM).

Females of doubtful identity.

Simbu: 27 females "Papua New Guinea: Supa Haia, 1023m, 10.ix.2002, K.Sagata (WB1)" (ZSM); these females are a mixture of three species: E. bacchusi , E. warasera , and E. haia . 5 females "Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 600m, 12IX2002, Balke & Sagata, (PNG 003)" (ZSM); these females are a mixture of two species: E. bacchusi and E. warasera . 3 females "Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 500m, 12IX2002, Balke & Sagata, (PNG 005)" (ZSM); these females are a mixture of three species: E. bacchusi , E. warasera , and E. haia . Simbu/Eastern Highlands: 3 females "Papua New Guinea: Simbu/EHPr. Crater Mountain, Sera - Herowana, Wara Pima, 900 m, 15IX2002, Balke & Sagata (PNG 011)" (ZSM). 20 females "Papua New Guinea: Crater Mountain, Sera - Herowana, upper Oh River, 1200 m, 15IX2002, Balke & Sagata (PNG 012)" (ZSM). 4 females "Papua New Guinea: Simbu/EHPr. Crater Mountain, Sera - Herowana, Jau river, 1000 m, 15IX2002, Balke & Sagata (PNG 015)" (ZSM). These females are a mixture of two species: E. bacchusi and E. warasera .

Diagnosis.

For complete description, see Balke (1998: 326). Beetle small to medium-sized: TL-H 3.05-3.9 mm, oblong-oval; dorsally uniformly reddish to dark brown or with paler head and sides of pronotum; shiny, with very fine to distinct punctation and usually weakly impressed microreticulation; pronotum with distinct lateral bead ( Fig. 16 ); male protarsomere 4 with anterolateral seta very long and thin, evenly curved, smaller than more laterally situated large seta; male protarsomere 5 ventrally with anterior band of more than 50 and posterior row of 8 relatively long setae ( Fig. 22D ); median lobe simple, evenly attenuated to broadly pointed apex in lateral and ventral views; paramere very slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin; proximal setae longer that subdistal, more distinct ( Fig. 22 A–C ).

Variability.

The species shows variability in size, colouration, how strongly impressed dorsal punctation and, more seldom, microreticulation, and slightly in shape of the apex of the median lobe ( Fig. 24 ).

Affinities.

From the species co-occurring in the same area ( E. craterensis Shaverdo & Balke, 2014, E. damantiensis (Balke, 1998), E. hintelmannae (Shaverdo, Sagata & Balke, 2005), E. sima , E. kobau sp. nov. and two species of the E. larsoni group), E. bacchusi can be distinguished by its reddish dorsal colouration and shape and setation of the median lobe and paramere. The most similar (in body size and form and colouration) to E. bacchusi are E. warasera sp. nov. and E. haia sp. nov., which occur with it. Only males of these species can be clearly separated by shape and setation of the median lobe and paramere; and therefore, dorsal setae of the paramere are important: in E. bacchusi , proximal setae longer that subdistal, more distinct. For the affinities within the group, see the “Key”.

Distribution.

Papua New Guinea: Madang, Simbu, Eastern Highlands, Morobe and Gulf Provinces ( Fig. 25 ). This is one of the most abundant species in the region.

Figures

Figure 24. Exocelina bacchusi (Balke, 1998), median lobe in lateral view A Madang, Adelbet Mt B Madang, Bismarck Range C Madang, Wilhelm Mt D Simbu, Crater Mt E EHL, Bena F Morobe, Kobau G Morobe, Yus H Gulf, Marawaka.

Figure 25. Map of the eastern part of New Guinea showing distribution of the species of the E. bacchusi group.

Figures 12 - 17. Habitus and colouration 12 Exocelina oiwa sp. nov. 13 E. akameku sp. nov. 14 E. erteldi (Balke, 1998) 15 E. oksibilensis sp. nov. 16 E. bacchusi (Balke, 1998) 17 E. bacchusi herzogensis sp. nov.

Figures 22, 23. 22 Exocelina bacchusi (Balke, 1998), paratype (Madang, Damanti) 23 E. bacchusi herzogensis ssp. nov. A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view.