SPERCHONTIDAE Thor, 1900
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publication ID |
https://doi.org/ 10.11646/zootaxa.2586.1.1 |
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persistent identifier |
https://treatment.plazi.org/id/5F029077-FF86-FFD6-FF22-FC21FE49FD09 |
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treatment provided by |
Felipe |
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scientific name |
SPERCHONTIDAE Thor, 1900 |
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Family SPERCHONTIDAE Thor, 1900 View in CoL View at ENA
Subfamily Sperchontinae Thor, 1900
Genus Sperchon Kramer, 1877
Sperchon (s. str.) brevirostris Koenike, 1895
New records: Bulgaria: Sofia: River Iskar above Samokov town , 1005 m a.s.l., 42 о 17'468"N 23 о 22'221"E, 19.viii.2006 Kenderov (0/2/0); Greece: Ipiros, Ioannina, stream Voidomatis, upstream of Aristi, 06.v.1997 Smit (0/1/1) .
Records from the study area: Bosnia and Hercegovina: Pešić (2003h); Montenegro: Pešić (1999, 2004a); Serbia: Ratajac & Rajaković (1992), Pešić (2003e); Macedonia: Schwoerbel (1963); Bulgaria: K. Viets (1926), Arndt (1943), Petrova (1971), Pešić (2006).
Remarks: New for Greece.
Habitat: Low- and middle-order streams.
Distribution: Western Palaearctic.
Sperchon (s. str.) glandulosus Koenike, 1886
New records: Croatia: Plitvice lakes area, stream Crna Rijeka, 24.vii.2007 Mičetić (0/1/0).
Records from the study area: Croatia: K. Viets (1936), Pešić (2002f); Bosnia and Hercegovina: Motaş et al. (1948); Macedonia: Pešić (2005); Bulgaria: Angelov (1964), Petrova (1971).
Habitat: Low- and middle-order streams.
Distribution: Central and Northern Europe, Balkan.
Sperchon (s. str.) longissimus K.Viets, 1920 Records from the study area: Bosnia and Hercegovina: Pešić (2003b)
Habitat: Crenobiont.
Distribution: England, Central and Southeastern Europe; Asia Minor.
Sperchon (s. str.) mutilus Koenike, 1895
New records: Montenegro: Bjelasica Mt., spring of Biogradska Rijeka , 42°51'N 19°40'E, 02.v.2003 Pešić (0/1/0); Andrijevica , Komovi Mt. , Štavna , 42°42'N 19°40'E, 27.v.2007 Pešić (0/1/0); Greece: GR17 Epiros: Pindos Metsovon , Pláma , beech forest spring, E-Hang, 1500 m a.s.l., QS 30 66, 23.v.1991 Gerecke (0/10/1); GR18 Pindos Metsovon , Pláma , spring 2 in beech forest, 1450 m a.s.l., QS 30 66, 23.v.1991 Gerecke (0/4/0); GR33 Makedonia, Vemon Pisoderion , spring 2 above village, 1500 m a.s.l., QN 47 72, 27.v.1991 Gerecke (0/ 2/0) GoogleMaps .
Records from the study area: Montenegro: Pešić (2004a); Macedonia: Schwoerbel (1963), Pešić (2005); Bulgaria: Petrova (1971), Pešić (2006).
Remarks: New for Greece.
Habitat: Crenobiont; preferably in cold rheohelocrenes.
Distribution: Central and Southern Europe.
Sperchon (s. str.) squamosus Kramer, 1879
Records from the study area: Bosnia and Hercegovina: Pešić (2003h); Montenegro: Pešić (1999); Bulgaria: K. Viets (1935b), Petrova (1971); Scheffler (1974).
Habitat: Rhithrobiont, crenophilous.
Distribution: Western Palaearctic.
Sperchon (s. str.) thienemanni Koenike, 1907
New records: Greece: Achaia : GR63 Erymanthos , Ano Vlassia, spring 2 at border of river Selinous, 1000 m a.s.l., 21°47'E 37°57'N, 27.v.1992 Gerecke (3/20/4); GR66 Erymanthos, Ano Vlassia, spring 1 northwest from pass Tripotama, 1700 m a.s.l., 21°44'E 37°57'N, 29.v.1992 Gerecke (0/2/0) GoogleMaps .
Records from the study area: Montenegro: Pešić (2004a); Macedonia: “ S. glandulosus “K. Viets (1936) (err.) - Gerecke (1991a); Bulgaria: “ S. rugosus ” Petrova (1971) , Pešić (2006).
Remarks: New for Greece.
Habitat: Crenophilous; with a preference for rheohelocrenes.
Distribution: Europe—exact distribution still unclear due to former confusion with S. glandulosus .
Sperchon (s. str.) vesnae Pesic, 2003
Records from the study area: Montenegro: Pesic (2003b, 2004a); Bulgaria: Pesic (2003b).
Habitat: Crenobiont.
Distribution: Balkan.
Sperchon (Hispidosperchon) clupeifer Piersig, 1896
New records: Montenegro: Bar, spring Škurta near village Dobre Vode , 12.v.2005 Pešić (1/0/0); Bar , Međuriječka Planina Mt. , stream, 15.iii.2007 Pešić (0/1/0); Macedonia : River Treska , 14.vii.1960, Petkovski (0/2/0); Serbia: Kuršumlija, River Banjska Reka , 07.viii.2003 Živić (0/1/0); Bulgaria: Sofia, River Iskar below Drujba , interstitial, 559 m a.s.l., 42 о 38'965"N 23 о 25'025"E, 15.i.2005 Kenderov (0/1/0); Eastern Rhodopes : Krumovgrad, stream near village Dolno Cerkoviste, 25.iv.2006 Pešić (0/4/0); tributary of Pecelarska River , 3 km from Asenovgrad, 23.iv.2006 Pešić (0/4/0); Greece: GR7p Evia Island , the region of Ochi, stream Gefira Dimosaris, downstream between Kalergo and Kallianos village, 30.vi.2007 Pešić (1/0/0); Thessalia, stream Sarandáporos, at junction of road E90/ road to Drosopigi, 05.v.1997 Smit (0/2/0); GR28 Makedonia, Vemon, affluent of river Aliakmon between Gauros and Aposkepos, 450 m a.s.l., 40°39'N 21°11'E, 26.v.1991 Gerecke (0/2/1) GoogleMaps .
Records from the study area: Croatia: K. Viets (1936), Pešić (2002f); Bosnia and Hercegovina: Motaş et al. (1948); Montenegro: K. Viets (1936); Serbia: Živić et al. (2002), Pešić (2003e); Macedonia: Schwoerbel (1963); Bulgaria: Petrova (1971); Scheffler (1974); Greece: K. Viets (1950).
Habitat: Low to high order streams.
Distribution: Western Palaearctic.
Sperchon (Hispidosperchon) gr. denticulatus
New records: Bosnia and Hercegovina: River Lištica near Široki Brijeg, 43°21'N 17°37'E, 12.xi.2006, Pešić (0/22/2); Montenegro: Berane , spring near Monastir Đurđevi Stupovi , 42°51'N 19°51'E, 29.ix.2009 Pešić (0/1/1); Pljevlja , River Breznica near Vodice , 793 m a.s.l., 43°21'52.51''N 19°21'49.50''E, 30.iv.2005 Pešić (0/23/0); Mojkovac, stream Ljevok near village Ljevok, 849 m a.s.l., 42°59'N 19°26'E, 10.viii.2006 Pešić (0/1/0); Bar, Međuriječka Planina Mt. , stream, 15.iii.2007 Pešić (0/6/0); Plav, Alipašini springs, 924 m a.s.l., 42°32'59.75''N 19°49´33.73''E, 04.x.2009 Pešić (0/1/0); Serbia: Sokobanja, the spring (“Vrelo“) of Moravica river , 390 m a.s.l., 43°37'53.77''N 21°59'9.15''E, 24.ix.2009 Pešić (0/1/0); Greece: Makedonia, Kozani , stream Pramoritsa , E of Mórfi (E of Kozani), 05.v.1997 Smit (0/1/0); Ipiros : GR8 Preveza , stream Buná Zalongu 2 km above Mirsíni, 180 m a.s.l., QW 78 93, 20.v.1991 Gerecke (0/2/46); GR9 Preveza , Buná Zalongu , affluent 2 km above Mirsíni, 180 m a.s.l., QW 78 93, 20.v.1991 Gerecke (0/0/5) GoogleMaps .
Records from the study area: Croatia: K. Viets (1936), Pešić (2002f); Bosnia and Hercegovina: Pešić (2003h); Montenegro: Pešić (1999); Serbia: Pešić (2003e); Macedonia: K. Viets (1936); Bulgaria: K. Viets (1926), Arndt (1943), Petrova (1971).
Remark: At present, species of this group are recognizable only in the male sex. As several species have been confused until recently ( Di Sabatino et al. 2009), all former records from the Balkan area are questionable. New for Greece.
Habitat: Low- to high-order streams.
Sperchon (Hispidosperchon) hibernicus Halbert, 1944
Records from the study area: Montenegro: Pešić (1999); Bulgaria: “ Sperchon monstruosus “ Pešić (2006) . Habitat: Middle order streams.
Distribution: British Isles, Central and southern Europe.
Sperchon (Hispidosperchon) hispidus Koenike, 1895
New records: Montenegro: River Morača near Međurječje , 186 m a.s.l., 42°43'05,43''N 19°22'06,80''E, 30.viii.1990 Pešić (0/1/0); Podgorica, River Morača near Duklja, 35 m a.s.l., 42°27'56,53''N 19°15'53,66''E, 10.vii.1999 Pešić (1/0/0); Danilovgrad, River Zeta near Spuž, 41 m a.s.l., 42°30'39,63''N 19°11'52,99''E, 08.vii.2001 Pešić (0/1/0); Podgorica, River Cijevna near village Dinoša, 97m a.s.l., 42°23'59,48''N 19°21'56,26''E, 01.vii.2000 Pešić (0/1/0); Štitarička River , between Mojkovac and Kolašin, 42 о 55'N 19 о 33'E, 23.vii.2001 Pešić (1/0/0); Bar, stream above Stari Bar, 42 о 05'59''N 19 о 08'30''E, 19.v.2004 Pešić (0/1/0); Komovi Mt. , River Opasanica near village Opasanica, 1139 m a.s.l., 42°40'36,07''N 19°32'52,38''E, 12.viii.1999 Pešić (1/0/0); Serbia: Niš, river Nišava , 01.viii.2003 Živić (0/2/0); Bulgaria: Sofia: River Iskar above Samokov town , 1005 m a.s.l., 42 о 17'468"N 23 о 22'221"E, 16.viii.2005 Kenderov (3/0); ibid., 26.xi.2005 Kenderov (13/1); Iskar River near Dragoshinovo village , 19.viii.2004 Kenderov (0/1/0); Cherni Iskar River , 1012 m a.s.l., 42 о 17'123''N 23 о 31'997''E, 19.v.2004 Kenderov (1/1/0); ibid., 16.viii.2005 Kenderov (1/3/0); Greece: Lesbos, stream Evergetoulas, 02.v.2000 Smit (3/6/0); Epiros: Ioannina, stream at junction of road E90 (Ioannina-Kozani) and road to Konitsa, 05.v.1997 Smit (91/17); GR4 Thesprotia, River Thyamis below Neráïda, 100 m a.s.l., QR 65 37, 18.v.1991 Gerecke (1/0/0); GR6 Thesprotia, Glyky, affluent of river Acheron from Frosíni, 150 m a.s.l., QR 82 15, 19.v.1991 Gerecke (1/0/0); GR12 Xerovuni, Plaka, River Arachthos below Agnanta, 200 m a.s.l., QS 15 24, 21.v.1991 Gerecke (1/5/5); Makedonia: GR44 Nestos- Delta, river course NE Chrysoupolis, 1 m a.s.l., RL 42 77, 30.v.1991 Gerecke (1/10/1); GR48 Vrondús, Sidirókastron, Krousovitis Potamos, 180 m a.s.l., QK 32 09, 02.vi.1991 Gerecke (3/3/1); Achaia: GR59 Erymanthos, Ano Vlassia, upper course of river Selinous , 1000 m a.s.l., 21°47'E 37°57'N, 25.v.1992 Gerecke (1/ 0/0); GR74 Aroánia, Kato Zachlorou, River Vouraikos near railroad station, 400 m a.s.l., 22°14'E 38°10'N, 05.vi.1992 Gerecke (3/6/26) GoogleMaps .
Records from the study area: Montenegro: Pešić (1999); Serbia: Pešić (2003d, e); Macedonia: Schwoerbel (1963); Bulgaria: Pešić (2006).
Remarks: New for Greece.
Habitat: Middle- and high-order streams.
Distribution: Western Palaearctic.
Sperchon (Hispidosperchon) papillosus Thor, 1901
New records: Greece: Evia Island: GR6p the region of Ochi, Agios Dimitrios Gorge, Porfyras Ravine stream, 30.vi.2007 Pešić (2/2/0); GR7p the region of Ochi, stream Gefira Dimosaris, downstream between Kalergo and Kallianos village, 30.vi.2007 Pešić (1/2/1); GR24p Sparta, stream Kelafina, a tributary of Evrotas river, 03.vii.2007 Pešić (0/3/0); Crete, stream E of Filaki, 27.iv.1987 Smit (2/0); Lesbos: stream Salantas, 04.v.2000 Smit (0/5/0); stream Priona, S of Neochori, 04.v.2000 Smit (0/1/0); Epiros, Ioannina, stream at junction of road E90 (Ioannina-Kozani) and road to Kanitsa, 05.v.1997 Smit (0/1/0); GR37 Makedonia, Chalkidiki, Chlomón Oros, Paleokastron, river Vatonia, 550 m a.s.l., QN 25 22, 28.v.1991 Gerecke (0/1/8).
Records from the study area: Macedonia: K. Viets (1936), Pešić (2005); Bulgaria: Pešić (2006).
Remarks: New for Greece.
Habitat: Low- and middle-order streams.
Distribution: Europe, Asia Minor, Iran.
Sperchon (Hispidosperchon) pelopeius Pešić & Gerecke sp. nov.
( Figs. 1–3 View FIGURES 1 View FIGURE 2 View FIGURE 3 )
Material examined. Holotype ( NHMB): female, dissected and slide-mounted in Hoyer's fluid. Greece, GR70 Achaia , Aroánia , Planiteron; rheocrene near Mazi, 1200 m a.s.l., 22°17'E 37°54'N, 02.vi.1992 Gerecke. Paratypes: (1/6/0), same data as holotype, one male of them dissected and slide-mounted GoogleMaps .
Further records. Achaia: Aroánia, GR73 Kato Zachlorou, spring in the Vouraikos valley, 500 m a.s.l., 38°10'N 22°14'E, 05.vi.1992 Gerecke (1/1/1); GR62 Erymanthos, Ano Vlassia, stream NW chapel, 1200 m a.s.l., 37°58'N 21°45'E, 26.v.1992 Gerecke (6/1/1); GR63 Erymanthos, Ano Vlassia, spring 2 at border of river Selinous, 1000 m a.s.l., 37°57'N 21°47'E, 27.v.1992 Gerecke (0/1/0).
Diagnosis. Integument ventrocaudally areolate, dorsally papillate. Dorsum with eight muscle attachment plates, two of which larger and including postocularia, the remaining in size not exceeding glandular diameter ( Fig. 1A View FIGURES 1 ). Cx-I+II medially not fused; Cx-III medially bearing Cxgl-4. Excretory pore smooth. Capitulum ( Figs 2F–G View FIGURE 2 ) with short rostrum; P-4 as long as, or only slightly longer than P-3, P-5 short, with strong distal claws; Leg ambulacrum with claw blade and dorsal and ventral clawlets.
Description
Integument ventrocaudally with fine denticles arranged in a hexagonal pattern; these denticles laterally and caudally inflating to form rounded papillae; these papillae regularly-arranged, nowhere fused to form lines or irregular structures, completely covering the dorsal surface. Dorsum with eight muscle attachment plates, following the terminology of Bader & Spasgozarian (1980) from anterior to posterior: medially, (1) small, round praefrontalia, (2) rather large plates consisting of fused postfrontalia, postocularia and dc-1, (3) very small dc-2 located close to each other, (4) larger and more distanced dc-3, (5) dc-4, smaller, but more distanced from each other than dc-3, and, laterally, (6–8) small, round platelets dl-1-3 located at the levels of interspace dc-1/2, dc-3, and interspace dc-3/4, respectively ( Fig. 1A View FIGURES 1 ). Ventral view ( Figs. 1B View FIGURES 1 , 2A View FIGURE 2 ): Cx-I+II medially close to each other, but not fused. Cx-III with a medial glandular opening (Cxgl-4). Ac-1-2 longish, Ac-3 roundish. An unpaired postgenital sclerite well distanced from genital field; other ventrocaudal sclerites absent, although muscules attached to cuticle on their places distinct. Excretory pore smooth. Capitulum ( Figs. 2F–G View FIGURE 2 ) with short rostrum; palps ( Figs. 1C–D View FIGURES 1 , 2D–E View FIGURE 2 ) normally held in a spread position (with their medial surface widely visible in dorsal view); P-1 without dorsal seta; P-2 with about 15 short and stout setae, all restricted to the dorsal part, distoventrally with a long projection at its tip bearing two fine setae, one long and one short; P-3 with smoothly convex ventral margin, bearing about 10 fine setae and several short hairs, all restricted to the dorsal part; P-4 relatively short, with the two ventral peg setae close to each other, the proximal one on a distinct elevation in the proximal part, the distal one minor and on a less developed thickening located slightly anterior to the centre of the segment; P-5 short, with strong distal claws; Leg segments slender, III-/IV-L with a few short, not plumose dorsal setae; ambulacrum with claw blade slightly protruding, bearing a long dorsal and a short ventral clawlet.
Female (holotype, in parentheses specimen from GR73): Idiosoma (ventral view: Fig. 1B View FIGURES 1 ) L/W 1000 (894)/856 (625). Distance between anterior end of Cx-I and posterior end of Cx-IV, 469 (403). Pregenital sclerite forming an obtuse triangle; genital valves L 167 (140), W 56 (50); L Ac-1-3 58 (42), 59 (46), 47 (35). Capitulum ( Fig. 2G View FIGURE 2 ) L 266 (215); chelicera L 281 (236), basal segment L 206 (162), claw L 69 (69), ratio chelicerae basal segment/claw L 3.0 (2.4); palp ( Figs. 1C–D View FIGURES 1 ) total L 598 (513), dL: P-1, 25 (23); P-2, 147 (124); P-3, 188 (156); P-4, 188 (164); P-5, 50 (46); %L: P-1, 4.2 (4.5); P-2, 24.6 (24.2); P-3, 31.4 (30.4), P-4, 31.4 (32.0); P-5, 8.4 (9.0); P-2/P-4 ratio 0.78 (0.76), P-3 and P-4 similar in L. Leg segments L: I-L-1-6: 67 (45), 91 (53), 117 (88), 181 (134), 169 (131), 156 (128); IV-L-1-6: 119 (94), 138 (103), 159 (116), 275 (219), 255 (200), 200 (153).
Male (paratype, in parentheses specimen from GR73): Idiosoma L/W 706 (556)/531 (431). Distance between anterior end of Cx-I and posterior end of Cx-IV, 370 (325). Pregenital sclerite forming an acute triangle; genital valves L 132 (118), W 36 (38); L Ac 1-3 47 (41), 45 (39), 36 (31). Ejaculatory complex ( Fig. 2B View FIGURE 2 ) L 164. Capitulum ( Fig. 2F View FIGURE 2 ) L 194 (168); chelicera ( Fig. 7 View FIGURE 7 ) L 198 (180), basal segment L 141, claw L 56, ratio chelicerae basal segment/claw L 2.5; palp ( Figs. 2D–E View FIGURE 2 ) total L 440 (388), dL: P-1, 21 (15); P-2, 100 (86); P-3, 129 (117); P-4, 148 (132); P-5, 42 (38); %L: P-1, 4.8 (3.9); P-2, 22.7 (22.2); P-3, 29.3 (30.2); P-4, 33.6 (34.0); P-5, 9.6 (9.8); P-2/P-4 ratio 0.68 (0.65), P-4 slightly longer than P-3. Leg segments L: I-L-2-6: 72 (53), 94 (71), 144 (116), 136 (116), 131 (109); IV-L-1-6: 106 (81), 103 (88), 120 (94), 216 (175), 206 (169), 169 (144).
Deutonymph (from GR73): Idiosoma L/W 438/338. Distance between anterior end of Cx-I and posterior end of Cx-IV 233. Provisional genital field ( Fig. 3A View FIGURE 3 ) with two pairs of Ac, L/W 54/61. Capitulum L 118, chelicera L 124; palp ( Fig. 3B View FIGURE 3 ) total L 296, dL (%L in parentheses): P-1, 17 (5.7); P-2, 68 (23.0); P-3, 89 (30.1); P-4, 95 (32.1); P-5, 27 (9.1).
Etymology. Named after the country of the type locality.
Discussion. In the shape of the palp combining a P-2 bearing a finger-like ventrodistal extension and a rather stout P-4 with the ventral peg setae strong, located close to each other on elevations near the segment centre, Sperchon pelopeius sp. nov. differs from all species known in the subgenus except for the rather similar Sperchon afzalipouri Bader & Sepasgozarian, 1980 , described from Iran (Semnan Province, Bader & Sepasgozarian 1980). This species differs in the absence of glandularia on Cx-III; a more elongated, fingershaped distoventral projection of P-2 and a stouter P-4, in both sexes being shorter than P-3.
Habitat: Crenobiont.
Distribution. Only found in rheocrene springs in the N Peloponnese.
Sperchon (Hispidosperchon) setiger Thor, 1898
New records: Croatia: Plitvice lakes area, Labudovac barrier, 28.vii.2007 Mičetić (0/1/0); Plitvice lakes area, Kozjak, 30.v.2007 Mičetić (0/1/0); Greece: Lesbos, stream Krioneri (stream and pools), 02.v.2000 Smit (1/1/4).
Records from the study area: Croatia: K. Viets (1936), Pešić (2002f); Serbia: Đorđević (1903), Pešić (2003e); Macedonia: Schwoerbel (1963); Bulgaria: Petrova (1971).
Remarks: New for Greece.
Habitat: Middle-order streams; often also in pool habitats.
Distribution: Palaearctic.
Sperchon (Hispidosperchon) violaceus Walter, 1944
Records from the study area: Montenegro: Pešić (1999, 2004a).
Habitat: In montane middle-order streams and springs.
Distribution: Western Palaearctic.
Genus Sperchonopsis Piersig, 1896
Sperchonopsis procera Láska, 1965
Records from the study area: Montenegro: Di Sabatino et al. (2009).
Habitat: Middle-order streams, hyporheobiont.
Distribution: Eastern Europe, only known from three sites in Slovakia, Poland and Montenegro.
Sperchonopsis verrucosa (Protz, 1896)
New records: Croatia: Plitvice lakes area, stream Crna Rijeka , 24.vii.2007 Mičetić (1/1/0), ibid., 27.viii.2007 Mičetić (2/2/0); Plitvice lakes area, Bijela Rijeka stream, 21.ix.2007 Mičetić (0/1/0); Montenegro: River Cijevna near village Dinoša (Podgorica), interstitial, 97 m a.s.l., 42°23'59,48''N 19°21'56,26''E, 25.viii.1999 Pešić (0/0/1); Komovi Mt. , small stream near village Opasanica, 1180 m a.s.l., 42°40'52,63''N 19°33'01,64''E, 09.xi.2001 Pešić (3/1/0); Pljevlja, River Breznica near Vodice, 793 m a.s.l., 43°21'52.51''N 19°21'49.50''E, 30.iv.2005 Pešić (1/2/0); Nikšić, River Vidrovanska rijeka near village Vidrovan, 657 m a.s.l., 42°51'N 18°56'E, 06.iv.2000 Pešić (0/0/1); Macedonia: River Bošava , 06.vii.1960, Petkovski (1/0/0); Greece: Makedonia: GR29 Vemon Pisoderion, affluent of river Ladopotamos , 1200 m a.s.l., 40°47'N 21°24'E, 26.v.1991 Gerecke (0/1/0); GR48 Vrondús, Sidirókastron, Krousovitis Potamos, 180 m a.s.l., QK 32 09, 02.vi.1991 Gerecke (0/1/0); Achaia: GR68 Aroania, Kato Klitoria, Aroanias river , Ag. Vlasios, 500 m a.s.l., 37°52'N 22°12'E, 31.v.1999 Gerecke (1/1/1); GR73 Aroánia, Kato Zachlorou, spring in the Vouraikos valley , 500 m a.s.l., 38°10´N 22°14´E, 05.vi.1992 Gerecke (0/2/0) GoogleMaps .
Records from the study area: Croatia: K. Viets (1936), Pešić (2002f); Bosnia and Hercegovina: Motaş et al. (1948); Montenegro: Pešić (1999); Serbia: Pešić (2003e); Macedonia: K. Viets (1936), Georgiev (1957), Schwoerbel (1963), Pešić (2005); Bulgaria: “ Pseudosperchon verrucosus ” K. Viets (1926) , Petrova (1971), Scheffler (1974); Greece: K. Viets (1950).
Habitat: In low-order streams, often on muddy, fine sediment; occasionally in the hyporheic; rhithrobiont.
Distribution: Holarctic.
| NHMB |
Natural History Museum Bucharest |
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