Megagonoleon, Machado & Oswald, 2020

Machado, Renato Jose Pires & Oswald, John David, 2020, Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae), Zootaxa 4796 (1), pp. 1-322 : 185-186

publication ID	https://doi.org/ 10.11646/zootaxa.4796.1.1
publication LSID	lsid:zoobank.org:pub:66DD1FEB-6BDE-4AEB-8A7B-96594371E9C5
persistent identifier	https://treatment.plazi.org/id/5F2387E7-70BB-FF1B-FF5F-FC84FA741C22
treatment provided by	Plazi
scientific name	Megagonoleon
status

Treatment

Key to species of Megagonoleon

1 Profemur <3.5x length of procoxa; pronotum not longer than wide (wider than long or as wide as long) ( Fig. 93b View FIGURE 93 )....... 4

- Profemur ≥ 3.5x length of procoxa; pronotum longer than wide ( Figs. 97b View FIGURE 97 , 104b View FIGURE 104 )......... ( cahillensis species group) … 2

2 (1’) Hind wing without a dark longitudinal stripe on posterodistal margin ( Fig. 97c View FIGURE 97 ); pronotum with posterior margin as wide as anterior margin ( Fig. 97b View FIGURE 97 )............................................................................. 3

- Hind wing with a dark longitudinal stripe on posterodistal margin ( Fig. 104c View FIGURE 104 ); pronotum with posterior margin much wider than anterior margin ( Fig. 104b View FIGURE 104 )................................................................ M. lesouefi

3 (2) Frons mostly black, but ventral margin pale ( Fig. 97a View FIGURE 97 ); hind wing with a large rhegmal infuscation ( Fig. 97c View FIGURE 97 ); female ectoproct without cavisetae ( Fig. 98d View FIGURE 98 ); male paramere apex strongly bent (ventral view) ( Fig. 98c View FIGURE 98 )............. M. cahillensis

- Frons mostly pale, but area between antennae black ( Fig. 111a View FIGURE 111 ); hind wing with a small rhegmal infuscation ( Fig. 111d View FIGURE 111 ); female ectoproct with cavisetae ( Fig. 112e View FIGURE 112 ); male paramere apex evenly curved (ventral view) ( Fig. 112d View FIGURE 112 ).................................................................................................... M. newi sp. nov.

4 (1) Profemur sense hair absent; tibial spurs absent or short (not reaching T1 apex; but reaching T1 apex in M. rebellis ); male gonarcus/mediuncus complex not semi-triangular in outline (dorsal view); male paramere well developed ( Fig. 118b View FIGURE 118 )........................................................................... ( pentagrammus species group) … 5

- Profemur sense hair present; tibial spurs present and long (reaching T1 apex; but absent in M. drysdalensis ); male gonarcus/ mediuncus complex semi-triangular in outline (dorsal view); male paramere absent or reduced (very thin, straight and entirely or mostly hidden beneath the gonarcus/mediuncus complex, in dorsal view) ( Fig.134c View FIGURE 134 ).....( stigmatus species group) … 15

5 (4) Tibial spurs absent................................................................................... 6

- Tibial spurs present................................................................................. 12

6 (5) Forewing with dark bands or scattered dark marks ( Fig. 117c View FIGURE 117 ); female anterior gonapophysis absent; male parameres not fused medially ( Fig. 118c View FIGURE 118 )............................................................................. 7

- Forewing hyaline ( Fig. 115c View FIGURE 115 ); female anterior gonapophysis present ( Fig. 116d View FIGURE 116 ); male parameres fused medially ( Fig. 116c View FIGURE 116 )........................................................................... M. oombulgurriensis sp. nov.

7 (6) Hind wing with one or more dark longitudinal lines ( Fig. 138c View FIGURE 138 )............................................... 8

- Hind wing without dark longitudinal lines ( Fig. 117c View FIGURE 117 )...................................................... 11

8 (7) Forewing mediocubital area not entirely brown ( Fig. 138c View FIGURE 138 )................................................... 9

- Forewing mediocubital area entirely brown ( Fig. 108c View FIGURE 108 )........................................... M. lulinguensis

9 (8) Clypeus entirely pale ( Fig. 131a View FIGURE 131 ); forewing crossveins not dark margined ( Fig. 131c View FIGURE 131 )............................. 10

- Clypeus pale with two brown spots ( Fig. 138a View FIGURE 138 ); forewing crossveins mostly dark margined ( Fig. 138c View FIGURE 138 )........ M. tindalei

10 (9) Frons mostly black ( Fig. 131a View FIGURE 131 ); male paramere acute posteromedially (lateral view) ( Fig. 132c View FIGURE 132 ).............. M. sarahae

- Frons mostly pale ( Fig. 99a View FIGURE 99 ); male paramere rounded posteromedially (lateral view) ( Fig. 100b View FIGURE 100 )............ M. cardaleae

11 (7’) Clypeus pale with two dark spots ( Fig. 113a View FIGURE 113 ); forewing with large dark marks ( Fig. 113c View FIGURE 113 )................ M. nigrescens

- Clypeus entirely pale ( Fig. 117a View FIGURE 117 ); forewing with no large dark marks ( Fig. 117c View FIGURE 117 )..................... M. pentagrammus

12 (5’) Wings relatively broad, ventral margin strongly convex below rehgma ( Fig. 106c View FIGURE 106 ); female ectoproct without cavisetae, lateral gonapophysis smaller than ectoproct, digitiform membranous process very long ( Fig. 107e View FIGURE 107 )........................ 13

- Wings relatively narrow, ventral margin weakly convex below rehgma ( Fig. 95c View FIGURE 95 ); female ectoproct with cavisetae, lateral gonapophysis about same size as ectoproct, digitiform membranous process absent ( Fig. 96e View FIGURE 96 )...................... 14

13 (12) Body mostly pale; wings hyaline ( Fig. 135c View FIGURE 135 ); female pregenital plate small and semi-elliptical ( Fig. 136f View FIGURE 136 ); male paramere with anterior region broad (ventral view) ( Fig. 136d View FIGURE 136 )................................................ M. tillyardi View in CoL

- Body mostly dark; wings with brown marks on distal half ( Fig. 106c View FIGURE 106 ); female pregenital plate large and kite shaped ( Fig. 107f View FIGURE 107 ); male paramere with anterior region narrow (ventral view) ( Fig. 107c View FIGURE 107 )................... M. longidigitus sp. nov.

14 (12’) Tibial spurs short (reaching half T1 length); forewing heavily marked, most crossveins brown margined ( Fig. 95c View FIGURE 95 ); male paramere apex acute (posterior view) ( Fig. 96c View FIGURE 96 ); female pregenital plate small, narrow, with medial region acute ( Fig. 96f View FIGURE 96 )................................................................................ M. breviplectron sp. nov.

- Tibial spurs long (reaching T1 apex); forewing not heavily marked, with small marks in mediocubital area and gradate crossveins ( Fig. 126c View FIGURE 126 ); male paramere apex rounded (posterior view) ( Fig. 127c View FIGURE 127 ); female pregenital plate large, rhomboidal ( Fig. 127e View FIGURE 127 )............................................................................... M. rebellis sp. nov.

15 (4’) Tibial spurs present................................................................................. 16

- Tibial spurs absent....................................................................... M. drysdalensis

16 (15) Body mostly black, with small pale areas................................................................ 17

- Body mostly pale to orange, with small black areas........................................................ 18

17 (16) Hind wing with some radial crossveins margined, rhegmal and stigmal marks about same size ( Fig. 124c View FIGURE 124 ); clypeus mostly dark ( Fig. 124a View FIGURE 124 )............................................................................. M. radialis

- Hind wing radial crossveins not margined, stigmal mark much larger than rhegmal mark ( Fig. 133c View FIGURE 133 ); clypeus pale ( Fig. 133a View FIGURE 133 ).................................................................................... M. stigmatus

18 (16’) Forewing subcostal and mediocubital areas hyaline or with only a few crossveins narrowly margined ( Fig. 129c View FIGURE 129 ); female terminalia: ventral membrane without thickened setae ( Fig. 130f View FIGURE 130 )............................................. 19

- Forewing subcostal and mediocubital areas with a series of dark rounded marks ( Fig. 122c View FIGURE 122 ); female terminalia: ventral membrane with numerous long thickened setae ( Fig. 123f View FIGURE 123 )....................................... M. punctatus sp. nov.

19 (18) Female pregenital plate conically invaginated ( Figs. 130 View FIGURE 130 e–f); male gonarcus/mediuncus complex elongate, completely covering paramere in dorsal view ( Fig. 130c View FIGURE 130 )................................................................. 20

- Female pregenital plate U-shaped ( Fig. 121f View FIGURE 121 ); male gonarcus/mediuncus complex not elongate, not covering anterior margin of paramere in dorsal view ( Fig. 121b View FIGURE 121 ).................................................................. 21

20 (19) Interantennal area mostly black ( Fig. 129a View FIGURE 129 ); male paramere not visible in lateral view ( Fig. 130b View FIGURE 130 ); female posterior gonapophysis absent; female 7 th sternite: posterior margin bearing two setose plates ( Fig. 130g View FIGURE 130 )................... M. roseipennis

- Interantennal area mostly pale ( Fig. 90a View FIGURE 90 ); male paramere visible in lateral view ( Fig. 91c View FIGURE 91 ); female posterior gonapophysis present ( Fig. 91f View FIGURE 91 ); 7 th sternite: posterior margin not bearing setose plates............................... M. annulatus

21 (19’) Supra-antennal area mostly pale to orange ( Fig. 93a View FIGURE 93 ); profemur and basal abdominal segments with numerous long white setae....................................................................................... M. banksi

- Supra-antennal area mostly black ( Fig. 120a View FIGURE 120 ); profemur and basal abdominal segments with short white setae and a few long black setae............................................................................. M. pingrupensis