Toxidium lunatum, Löbl, 2012

Löbl, Ivan, 2012, On a collection of Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) from West Malaysia, Acta Entomologica Musei Nationalis Pragae 52 (1), pp. 173-184 : 182

Toxidium lunatum sp. nov.

( Fig. 15 )

Type material. HOLOTYPE: ♂, ‘ 1.–3.IV.1990 Malaysia Cameron Highl. Gn. Beremban leg. A. Riedel’ ( MHNG). PARATYPES: 1 ♀, ‘ 20.IV.1990 Malaysia Cameron Highl. Gn. Brinchang leg. A. Riedel’ ( MHNG); 1 ♀, ‘ MALAYSIA, Pahang Cameron Highlands Tanah Rata vill. env. Gunung Jasat [Mt.] ; 1470–1705 m 04°28.4–7’ N, 101°21.6–22.1’ E Jiří Hájek leg. 18.iv.–10.v.2009 ’ ( NMPC); 2 ♀♀, ‘W. Malaysia Pahang Cameron Highlands 12.–15.ii.1998 Tanah Rata , Gn. Jasar lgt. S. Becvar’ ( MHNG); 1 ♀, ‘ MALAYSIA-W, Pahang, 30 km SE of Ipoh, 1500m, Banjaran Titi Wangsa, Tanah Rata , 14–15.iii. 2002, P. Černohorský leg.’ ( NMHW).

Description. Length 2.10–2.50 mm, width 1.15–1.27 mm. Head and most of body very dark, blackish with reddish shine or black. Elytra each with C-shaped, reddish fascia open laterally, narrowed along suture. Femora and tibiae dark reddish-brown. Apical abdominal segments, tarsi and antennomeres I and II, eventually also III and IV light, ochraceous, following antennomeres darkened. Length ration of antennomeres as: II 13: III 11: IV 12: V 16: VI 13: VII 17: VIII 13: IX 16: X 14: XI 19 (holotype). Segments III and IV even, very narrow, segments V and VI slightly wider. Pronotal punctation very fine, consisting of comparatively well delimited punctures distinct at 25 times magnification. Tip of scutellum exposed. Elytra with sutural striae very shallow, variable in length, starting slightly posterior to basal fourth, or more apically, beyond middle of sutural length; basal striae present, very shallow, variably long, complete and joined to lateral striae, reaching almost level of scutellum or shortened and visible only in middle section of elytral base; lateral and epipleural striae converging in apical part only, lateral stria coarsely and densely punctate, discal punctation not forming rows, irregularly coarse and dense, most punctures clearly smaller than puncture intervals. Metathoracic wings fully developed. Mesoventrite lacking distinct median ridge, with two very shallow and short admesal striae or impressions. Metaventrite flattened in middle, appearing impunctate, most punctures hardly visible at 100 times magnification, few very fine punctures visible at lower magnification; submesocoxal line convex, with few distinct punctures; submesocoxal area 0.08 mm, about as long as half of shortest interval to metacoxa. Metepisternum flat, hardly narrowed apically, with suture straight, deep and wide, in particular at proximal end. Exposed abdominal segments with extremely fine punctation hardly visible at 100 times magnification, microsculpture apparently absent. Tibiae straight.

Male. Tarsomeres 1 to 3 of prolegs similar, slightly widened. Aedeagus ( Fig. 15 ) 0.58 mm long. Median lobe gradually narrowed apically, with apical process weakly inflexed and shorter than basal bulb. Parameres narrowed in middle, with apical lobe weakly sclerotized and setose apical margin. Internal sac with long rod followed by membranes very finely longitudinally striate.

Differential diagnosis. The colour pattern of elytra separates this species from other Asian congeners (see the key in LÖBL (1999)). Its aedeagus is similar to that in T. pubistylis Löbl, 1990 suggesting close relationships. The parameres in the latter species are distinctive, having an apical setose lobe, and the membranous structures of the internal sac are not longitudinally striate.

Etymology. The name is a Latin adjective, referring to the crescent shape of the reddish elytral band.

Figures

Figs. 13–15. 13–14 – Scaphisoma edentatum sp. nov.: 13 – aedeagus in lateral view, scale bar = 0.2 mm; 14 – internal sac in lateral view, scale bar = 0.1 mm. 15 – Toxidium lunatum sp. nov., aedeagus in dorsal view, scale bar = 0.2 mm.