Platyderus (Eremoderus) brunneus Karsch, 1881

Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice, 2022, Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini), Deutsche Entomologische Zeitschrift 69 (1), pp. 71-122 : 71

publication ID	https://dx.doi.org/10.3897/dez.69.83840
publication LSID	lsid:zoobank.org:pub:7DEB0587-A373-4B06-BAAD-CA6102EC6581
persistent identifier	https://treatment.plazi.org/id/5F6B14D7-3F4F-589A-B642-1AE0A0780643
treatment provided by	Deutsche Entomologische Zeitschrift by Pensoft
scientific name	Platyderus (Eremoderus) brunneus Karsch, 1881
status	stat. rev.

Treatment

15. Platyderus (Eremoderus) brunneus Karsch, 1881 View in CoL stat. rev.

Figs 3D, F View Figure 3 , 5H View Figure 5 , 6A, B View Figure 6 , 7I, J View Figure 7 , 9C, D View Figure 9 , 11B, C View Figure 11 , 12J, K View Figure 12 , 15A, B View Figure 15 , 18 View Figure 18

Diagnosis.

A species of Platyderus (Eremoderus) , with mesotarsomeres and metatarsomeres dorsally more or less flattened and grooved (Fig. 6A, B View Figure 6 ), cordate pronotum, with sides from widest point to base markedly (nominotypical subspecies) or barely concave (ssp. Platyderus brunneus ferrantei ) and posterior angles obtuse, with tips laterally more (nominotypical subspecies) or less protruding (ssp. Platyderus brunneus ferrantei ), and median lobe of aedeagus with basal bulb more slanting to shaft (Fig. 9C, D View Figure 9 ).

Redescription.

Habitus. Moderately to large-sized species (BL: 7.60-9.30 mm; BW: 2.70-3.40 mm), with elongate and moderately convex body (Fig. 3D, F View Figure 3 ). Measurements and ratios. See Table 3 View Table 3 . Color and lustre. Body, palpi, antennae and most of legs yellow-brown to reddish- brown, with head usually darker than pronotum and elytra and femora lighter than tibiae and tarsomeres. Integument slightly shiny (less shiny than in P. languidus ). Microsculpture and punctation. Pronotum with evident microreticulation, sculpticells regularly isodiametric. Elytral intervals, scutellum and lateral gutterwith distinct isodiametric sculpticells; basal margin with reduced microreticulation. Ventral surface with impressed slight transverse sculpticells or with sculpticells scarcely-visible. Head impunctate, usually smooth, clypeus and areas laterad of frontal furrows sometimes with fine longitudinal wrinkles. Surface of pronotum impunctate, largely smooth, only basal area medially behind posterior transverse impression longitudinally wrinkled, rarely apical part medially in front of anterior transverse impression also wrinkled; basal foveae and adjacent lateral areas smooth, smoother than those in P. languidus . Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled in whole, 2-5 finely wrinkled at sides. Head. About one-third as wide as pronotum (PW/HW= 1.28-1.41). Eye long, moderately convex. Labrum subrectangular, as long as clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, reduced to disappeared at sides. Frontal furrows small, rather shallow, punctiform. Paraorbital sulci straight, fine, backward surpassing posterior margin of eye, nearly reaching level of posterior supraorbital pore. Thorax. Pronotum one-tenth to one-fifth wider than long (PW/PL= 1.10-1.21), with widest point at second quarter. Anterior transverse impression absent, posterior transverse impression short, barely distinct. Sides convex medially and anteriorly, slightly concave to straight in base; anterior bead present laterally, lacking or present in medial eight to tenth; basal bead present laterally, reduced to absent in medial fifth to half. Metepisternum slightly longer than wide or nearly as long as wide, MA/MI= 0.87-1.00, with greater values occurring in specimens of ssp. Platyderus brunneus ferrantei . Elytra. Oblong, about one-and-a-half times to one and two thirds as long as wide (EL/EW= 1.52-1.72), two-and-a-half times or little more as long as pronotum (EL/PL= 2.46-2.72), and one and a third to a one-and-a-half times as wide as pronotum (EW/PW= 1.35-1.48), with widest point at medial third. Parascutellar striole and striae 1-7 very shallow to indistinct, stria 8 evident, clearly deeper than other striae (nominotypical form) or parascutellar striole and striae 1-7 distinct, slightly impressed, stria 8 somewhat deeper than other striae (ssp. Platyderus brunneus ferrantei ); parascutellar striole, if present, not joining stria 1; striae 1-8 superficially to moderately punctate; bases of striae 1-7 more or less reduced, not reaching basal bead (nominotypical form) or distinct, reaching basal bead (ssp. Platyderus brunneus ferrantei ). Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 or 17 punctures on each side. Legs. Posterior side of profemur with one or two setae in basal third and one or two in medial third. Mesofemur with 4-7 setiferous punctures on anterior side ventrally (Fig. 5H View Figure 5 ). Anterior side of metafemur ventrally with a few long setae, one in basal third and one to three in apical half. Male genitalia. Urite IX suboval, with proximal margin slightly asymmetrical, insignificantly turned to left (Fig. 7I, J View Figure 7 ). Median lobe of aedeagus in lateral view slender, with narrow basal bulb, long shaft and short, straight apex (Fig. 9C, D View Figure 9 ); median lobe in ventral view straight, 3.4-3.5 times longer than wide (Fig. 11B, C View Figure 11 ); apical lamella (dorsal view) short, scarcely asymmetrical, rounded at tip, with right side straight and left side slightly concave. Internal sac in lateral view (Fig. 9C, D View Figure 9 ) with ventral sclerite long, broadened and rounded distally; same in ventral view (Fig. 11B, C View Figure 11 ), with dorsal sclerite large, reticulate, with left-sided and right-sided protuberances distinct, and ventral sclerite narrow and distally very slightly curved to left. Right paramere as on Fig. 12J, K View Figure 12 . Female genitalia (Fig. 15A, B View Figure 15 ). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.

Notes.

Due to minor morphological differences, the populations from the lower course of Nile River and Southwest Israel are treated as a distinct subspecies of P. brunneus (see Fig. 18 View Figure 18 ). Moreover, no verifiable records of the species are so far known from the vast area between Misrata District (Libya) and Cairo Governorate (Egypt).