Senaspis dentipes (Macquart, 1842)

Meyer, Marc De, Goergen, Georg & Jordaens, Kurt, 2020, Taxonomic revision of the Afrotropical hover fly genus Senaspis Macquart (Diptera, Syrphidae), ZooKeys 1003, pp. 83-160 : 83

Senaspis dentipes (Macquart, 1842) Figs 1 , 12-15 , 52 , 64 , 80 , 91 , 102

Eristalis dentipes Macquart, 1842: 97.

Helophilus aesacus Walker, 1849: 609. Syn. by Smith and Vockeroth (1980).

Plagiocera maculipennis Loew, 1858: 381. Syn. by Austen (1909).

Eristalis latevittatus Bigot, 1858: 365. Syn. by Kertész (1910).

Protylocera aesacus var. livida Bezzi, 1912: 421. Syn. nov.

Differential diagnosis.

A predominantly subshiny brownish species (Fig. 1 ), without distinct pilosity. The wing (Fig. 64 ) is largely hyaline but with a distinct dark brown medial macula. It can be differentiated from other Senaspis species with a distinct wing macula ( S. haemorrhoa , S. xanthorrhoea ) by the absence of orange pile on the abdominal terga and the presence of a single distinct ventral swelling in the apical fifth of the metafemur (Fig. 80 ) (two swellings in S. haemorrhoa and S. xanthorrhoea ).

Examined material.

Eristalis dentipes Macquart: Lectotype (hereby formally designated and published; see comments), male, "N° 1177. // Eristalis // Eristalis dentipes ." "272 // 97" “70” "MNHN, Paris // ED6399" “LECTOTYPE” "TYPE // Vockeroth ‘69” "LECTOTYPE // Eristalis // Eristalis dentipes Macq. // Desig. Thompson 1977" [MNHN].

Helophilus aesacus Walker: Holotype, male, "Holo- // type" “Type” " Helophilus // Helophilus aesacus // Wlk." "Sierra Leone" [NHMUK].

Eristalis latevittatus Bigot: Holotype, female, "2423 // 83" “TYPE” " Eristalis // Eristalis latevittatus ♀. // n.sp. J. Bigot. // Gabon. // Coll. Thomson." [MNHN].

Protylocera aesacus var. livida Bezzi: Lectotype (hereby designated), male "Is. Fernando Poo // Basile // 400-600 m.s.m. // VIII-IX.1901. L. Fea leg.;" “Typus” "SYNTYPUS // Protylocera // Protylocera aesacus var. // Protylocera livida Bezzi, 1912" " Protylocera // Protylocera aesacus Walker // v. Protylocera livida // det. Bezzi, 1912" "Museo Cvico // di Genova" " Protylocera // Protylocera aesacus // var. lívida n." “LECTOTYPUS” [MCSNG]. Paralectotype, female, "Is. Fernando Poo // Musola // 500-800 m.s.m. // I III.1902. L. Fea leg.;" "SYNTYPUS // Protylocera // Protylocera aesacus // var. lívida” " Protylocera // Protylocera aesacus Walker // v. Protylocera livida // det. Bezzi, 1912" "Museo Civico // di Genova" "PARA- // LECTOTYPUS" [MCSNG].

Other material.

Angola • 1♀; Congulu, Apr. 1934; K. Jordan leg.; NHMUK • 1♂; 7 mi W Gabela; 16-18 Mar. 1972; NHMUK. Benin • 1♀; Ahozon; Jun. 2005; G. Goergen leg.; IITA • 1♂; Calavi, IITA Station; 4 Apr. 1994; G. Goergen leg.; IITA • 1♀; same collection data as for preceding; 4 Aug. 1995 • 1♀; same collection data as for preceding; 5 Mar. 1997 • 1♂ 1♀; same collection data as for preceding; Jun. 2006 • 1♂; same collection data as for preceding; 18 Apr. 2007 • 4♂♂ 2♀♀; same collection data as for preceding; Jun. 2007 • 1♂; same collection data as for preceding; 16 Nov. 2011; K. Jordaens leg.; KMMA • 11♂♂ 3♀♀; same collection data as for preceding; 8 Dec. 2013 • 1♂; same collection data as for preceding; 16 Dec. 2013 • 3♀♀; Cotonou; 3 Jun. 1914; W.A. Lamborn leg.; NHMUK • 1♂; Cotonou; 24 Sep. 2002; G. Goergen leg.; IITA • 1♀; same collection data as for preceding; May 2006 • 1♀; same collection data as for preceding; 25 Apr. 2012 • 1♀; Dassa; Aug. 2007; G. Goergen leg.; IITA • 1♂; Lama forest; 26 Jul. 1995; G. Goergen leg.; IITA • 1♂; Lokoli; Feb. 2006; G. Goergen leg.; IITA • 1♂; Lokossa; Jan. 2005; G. Goergen leg.; IITA • 2♀♀; same collection data as for preceding; Oct. 2005 • 2♂♂; same collection data as for preceding; Jan. 2006 • 2♂♂; same collection data as for preceding; Mar. 2006 • 1♀; Niaouli; Jun. 2003; G. Goergen leg.; IITA • 1♀; Ouidah; Nov. 2004; G. Goergen leg.; IITA • 2♀♀; same collection data as for preceding; May 2006 • 1♂ 1♀; same collection data as for preceding; 23 Apr. 2009 • 1♂; Pénéssoulou; Jun. 2004; G. Goergen leg.; IITA • 1♂; Porto Novo; Aug. 2006; G. Goergen leg.; IITA • 1♂; Porto Novo; 27 Jan. 2016; K. Jordaens leg.; KMMA • 1♂; Tanougou Waterfalls; 20 Jan. 2020; G. Goergen leg.; IITA • 1♂; Togbin; Dec. 2005; G. Goergen leg.; IITA. Burundi • 1♂; Busoni, Kisenyi; 17 Dec. 1950; F. François leg.; KBIN • 2♂♂ 1♀; Kibira NP, Rwegura; 31 May 2018; E. Sinzinkayo leg.; OBPE • 1♀; same collection data as for preceding; 28 Nov. 2019 • 1♀; Kisenyi; May 1955; F.J. François leg.; KBIN • 1♀; Kitega; 22 Oct. 1950; F. François leg.; KBIN • 1♂; Lake Nyanza; 7 [?] 2013; L. Ndayikeza leg.; OPBE • 1♂; same collection data as for preceding; 23 [?] 2013 • 1♂; Nyakibande, Mumirwa; 27 Jan. 2018; E. Sinzinkayo leg.; OBPE • 1♀; same collection data as for preceding; 16 May 2018 • 1♂; same collection data as for preceding; 24 Apr. 2019 • 2♂♂ 1♀; same collection data as for preceding; 14 Jul. 2019 • 2♀♀; Nyamurembe, Bururi; 7 Mar. 1953; P. Basilewsky leg.; KMMA • 1♀; Rumonge; 1934; A. Lestrade leg.; KMMA • 1♀; Rumonge; 19-20 Jun. 1948; F. François leg.; KBIN • 1♀; same collection data as for preceding; 25 Nov. 1948 • 1♀; same collection data as for preceding; 25 Feb. 1949 • 1♀; same collection data as for preceding; 19 May 1952 • 1♀; Rumonge; 7 Mar. 1953; P. Basilewsky leg.; KMMA • 1♀ Usumbura [= Bujumbura]; G. Pierrard leg.; KMMA. Cameroon • 2♀♀; Bafut Forest, Nguenba; 29 Nov. 1967; de Miré leg.; MHNH • 1♂; Batanga; 24 Jun. 1911; A.I. Good leg.; CNC • 1♀; Ebogo; 9 Jul. 1999; G. Goergen leg.; IITA • 1♀; Kumba-Victoria Rd boundary post; 9 Nov. 1949; H. Oldroyd leg.; NHMUK • 1♀; Lolodorf; 28 Aug. 1915; A.I. Good leg.; CNC • 2♂♂; Mbalmayo; 3 Jul. 1998; G. Goergen leg.; IITA • 1♂; same collection data as for preceding; 2 Aug. 2000 • 1♀; Victoria [= Limbe]; Jun. -Aug. 1917; F.H. Fitz Roy leg.; NHMUK. Democratic republic of the Congo • 1♀; Abok; 1927; Ch. Scops leg.; KMMA • 2♀♀; Albertville [= Kalemie]; 14 Aug. 1953; J. Verbeke leg.; KBIN • 1♂; Bambesa; 30 Oct. 1933; J.V. Leroy leg.; KMMA • 1♂; Bambesa; 6 Jul. 1937; J. Vrydagh leg.; CNC • 1♀; same collection data as for preceding; 2 Oct. 1937; KMMA • 1♀; same collection data as for preceding; 24 Mar. 1939; KBIN • 1♀; same collection data as for preceding; 3 Jun. 1939; KBIN • 1♀; same collection data as for preceding; 11 Nov. 1940; KMMA • 1♀; Barumbu; Aug. 1925; J. Ghesquière leg.; KMMA • 1♂; Bafwasende; Nov. 1945; F. François leg.; KMMA • 1♂; Bezali; 3 Oct. 1968; P.M. Elsen leg.; KMMA • 1♀; Boma; R.F. Achille leg.; KMMA • 1♀; Boma; 17 Jun. 1915; Lang and Chapin leg.; KMMA • 1♀; Bomputu, Salonga; Jun. 1936; J. Ghesquière leg.; KBIN • 1♂; Bukama; 2 Apr. 1911; Bequaert leg.; KMMA • 1♀; Bumba; Dec. 1939-Jan. 1940; H. De Saeger leg.; KMMA • 1♀; Coquilhatville [= Mbandaka]; 1946; Ch. Scops leg.; KMMA • 1♂; between Deti and Ibambi; 5 Jul. 1931; F.R. Swift leg.; NHMUK • 1♀; Eala; Mar. 1935; A. Corbisier leg.; KMMA • 1♀; Eala; Mar. 1935; J. Ghesquière leg.; KBIN • 1♀; same collection data as for preceding; 12 Aug. 1935 • 1♀; same collection data as for preceding; 19 Nov. 1935 • 1♂; same collection data as for preceding; 12 Nov. 1936 • 1♀; Elisabethville [= Lubumbashi]; Nov. 1932; C.V. Hirschberg leg.; KBIN • 1♀; Elisabethville [= Lubumbashi]; Nov. 1933; M. Bequaert leg.; KBIN • 1♂; same collection data as for preceding; 22 Apr. 1934 • 1♀; same collection data as for preceding; 3 Mar. 1935 • 1♂; Elisabethville [= Lubumbashi]; 1-6 Sep. 1932; de Loose leg.; KMMA • 1♀; same collection data as for preceding; 24 Nov. 1932 • 1♀; Faradje; 30 Mar. 1912; Lang and Chapin leg.; KMMA • 1♀; same collection data as for preceding; Nov. 1912 • 1♀; Idjwi Island, Kivu; 8 Nov. 1952; J. Verbeke leg.; KBIN • 1♀; Jadotville [= Likasi]; J. Muller leg.; KBIN • 3♂♂; Kadjudju; Feb. 1932; G. Babault leg.; MHNH • 4♂♂ 4♀♀; same collection data as for preceding; May-Jun. 1932 • 2♂♂ 8♀♀; same collection data as for preceding; Aug. 1932 • 2♂♂; same collection data as for preceding; Sep. 1932 • 1♀; Kapanga; Nov. 1932; F.G. Overlaet leg.; KMMA • 2♂♂ 1♀; same collection data as for preceding; Dec. 1932 • 1♀; same collection data as for preceding; Mar. 1933 • 1♀; Kasongo; 10 Sep. 1959; P.L.G. Benoit leg.; KMMA • 1♀; Katoko-Kobe; 10-20 Jan. 1958; Segers leg.; KBIN • 1♀; Katompe; 12 Dec. 1923; M. Bequaert leg.; KMMA • 1♀; Kimuenza; Sep. 1962; M.J. Deheegher leg.; KMMA • 1♀; Kisantu; 1927; P. Vanderyst leg.; KMMA • 1♀; Komi, Sankuru; Dec. 1930; J. Ghesquière leg.; KMMA • 1♀; same collection data as for preceding; CNC • 2♂♂; Kwango; Panzi; 12 Feb. 1939; Mrs Bequaert leg.; KMMA • 1♀; Lubutu; 22 Jan. 1915; J. Bequaert leg.; KMMA • 1♀; Lukolela; 3 Jan. 1931;.P. Chapin leg.; AMNH • 1♂; same collection data as for preceding; 7 Jan. 1931 • 1♀; Luputa, Lomami; Jan. 1935; Bouvier leg.; KMMA • 1♀; same collection data as for preceding; Mar. 1935 • 1♂; same collection data as for preceding; May 1935 • 1♀; Mbandaka; 1 Feb. 1962; A.B. Stam leg.; KMMA • 1♀; Mboga; Jul. 1914; J. Bequaert leg.; KMMA • 1♀; Medje, Ituri; 9-15 Jul. 1910; Lang and Chapin leg.; AMNH • 1♂; Medje, Ituri; 1-7 Sep. 1910; Lang and Chapin leg.; KMMA • 1♂ 2♀♀; Moma, Equateur; Jun. 1925; J. Ghesquière leg.; KMMA • 1♀; N’Guli; 6-9 Oct. 1913; Rodhain leg.; KMMA • 1♀; Niangara, Uelé; Nov. 1910; Lang and Chapin leg.; KMMA • 1♀; Nzali, Ubangi; 3 Feb. 1932; H.J. Brédo leg.; KMMA • 2♀♀; Pawa, Uelé; 1938; A. Dubois leg.; KMMA • 1♀; Poko, Uelé; Aug. 1913; Lang and Chapin leg.; AMNH • 1♀; same collection data as for preceding; KMMA • 1♂; Pweto, Katanga; Oct. 1926; A. Bayet leg.; KMMA • 3♂♂ 1♀; Stanleyville [= Kisangani]; Mar. 1915; Lang and Chapin leg.; AMNH • 1♂; same collection data as for preceding; CNC • 3♂♂ 1♀; same collection data as for preceding; KMMA • 1♀; same collection data as for preceding; 6 Apr. 1915; CNC • 1♀; same collection data as for preceding; 7 Apr. 1915; KMMA • 1♀; same collection data as for preceding; 14 Apr. 1915 • 1♂; same collection data as for preceding; Apr. 1915 • 1♂; same collection data as for preceding; KBIN • 1♀; Tshibamba, Lulua; Mar. 1932; F.G. Overlaet leg.; KMMA • 4♀♀; Uvira, Kivu; 25 Aug. 1957; P. Basilewsky leg.; KMMA. Equatorial Guinea • 1♂; Fernando Po [= Bioko Island], Basile; Aug. 1901; L. Fea leg.; MCSNG • 1♂; Musola; Jan. 1902; L. Fea leg.; MCSNG • 1♀; same collection data as for preceding; Mar. 1902 • 1♀; Punta Frailes; Oct.-Nov. 1901; L. Fea leg.; MCSNG • 1♂ 1♀; San Benito [= Mbini]; 1885; Guiral leg.; MHNH. Ethiopia • 1♂; Arba-Minch, Lake Chano; 19 Oct. 2012; A. Pauly leg.; KMMA. Gabon • 1♂; Ipassa, route de Makokou; 2-17 May 1974; M. Donskoff and J. Le Breton leg.; MHNH. Ghana • 1♀; Accra; 1-20 Jul. 1988; M. Hauser leg.; CDFA • 1♀; Buroburo, Kumasi; 4 Jun. 1976; A.B. Stam leg.; KMMA • 1♀; same collection data as for preceding; 12 Jul. 1976 • 1♀; same collection data as for preceding; 17 Jul. 1976 • 1♀; same collection data as for preceding; 28 Jul. 1976 • 1♀; Cape Coast; Dec. 2004; G. Goergen leg.; IITA • 1♂; Obuasi, Ashanti; 8 Aug. 1906; W.M. Graham leg.; NHMUK • 2♂♂ 1♀; same collection data as for preceding; 25 Aug. 1907 • 1♀; Takoradi; 2-6 Jun. 1957; G.H. Heinrich leg.; CNC • 1♂; Tsito; 26 Jan. 1969; O.W. Richards leg.; NHMUK • 2♂♂ 1♀; Wati Waterfalls; Feb. 2003; G. Goergen leg.; IITA • 1♀; Worawora; Mar. 2005; G. Goergen leg.; IITA. Guinea Bissau • 1♂ 2♀♀; Bolama; Jun.-Dec. 1899; L. Fea leg.; MCSNG. ‘GUINEA’ • 1♀; Westermann leg.; MNB [labeled as type of Plagiocera maculipennis considered to originate from Guinea Bissau according Smith and Vockeroth (1980); cf. comments]. Ivory Coast • 1♀; Adiopodoumé; 4 Jul. 1948; Jouer leg.; MHNH • 1♂; Banco National Park; 23 Apr. 1989; J. Londt leg.; NMSA. Kenya • 1♀; Itieni Forest, Nyambene Hills; 16-30 Oct. 2011; R.S. Copeland leg.; ICIPE • 1♂; Kaimosi; Feb. 1949; V.G.L. van Someren leg.; NHMUK • 1♀; Lukusi River, Kakamega Forest; 5 Apr.-26 Jun. 2017; R.S. Copeland leg.; ICIPE • 1♂; Taveta Forest; Aug. 1947; V.G.L. van Someren leg.; NHMUK. Liberia • 1♂; Bendu, Robertsport; 11 Apr. 1943; F.M. Snyder leg.; AMNH • 1♀; Du River camp 3; 1926; J. Bequaert leg.; KMMA • 1♀; Kpaine; 17 Aug. 1953; W. Peters leg.; NHMUK • 1♂; Monrovia; 24 May 1957; G.H. Heinrich leg.; CNC. Malawi • 1♂; Mt Mlanje; 9 Oct. 1913; S.A. Neave leg.; NHMUK. Mozambique • 1♀; East of Mt Mlanje; 21 Nov. 1913; S.A. Neave leg.; NHMUK • 1♂; Kola Valley; 5 Apr. 1913; S.A. Neave leg.; NHMUK • 1♂; Rikatla; H.A. Junod leg.; NMSA. Nigeria • 1♂ 1♀; Afikpo; 2 Jun. 1910; J.J. Simpson leg.; NHMUK • 1♂; Gadau; Jul. 1938; Buxton and Lewis leg.; NHMUK • 2♂♂; Ibadan; 24 Jul. 1947; J.L. Gregory leg.; NHMUK • 1♀; same collection data as for preceding; Jun. 1951; CNC • 1♂; Ibadan, IITA station; 24 Jan. 1997; G. Goergen leg.; IITA • 1♀; same collection data as for preceding; 26 Jan. 1998 • 1♂ 1♀; Ile-Ife; 30 Oct. 1969; J.T. Medler leg.; CNC • 1♀; Jos; Oct.-Dec. 1965; E. Bot Gwong leg.; KMMA • 1♀; Kaduna; Aug. 1952; Nash leg.; NMSA • 2♀♀; Kaduna; 19 Sep. 1962; M.W. Service leg.; NHMUK • 1♀; Olokemeji, Ibadan; AMNH. Republic of the Congo • 1♀; Fernand-Vaz; Sep.-Oct. 1902; L. Fea leg.; MCSNG • 1♂; Haut Ivindu, Ogooué; 1906; J. Gravot leg.; MHNH • 1♂; Lambaréné, Ogooué; 1913; E. Ellenberger leg.; MHNH. Sierra Leone • 1♀; Bomaru; 8 Aug. 1912; J.J. Simpson leg.; NHMUK • 1♀; Daru; 31 Jul. 1912; J.J. Simpson leg.; NHMUK • 1♀; same collection data as for preceding; 18 Aug. 1912 • 1♀; Ka-Yima; 23 Jun. 1912; J.J. Simpson leg.; NHMUK • 1♀; Lungi; 14 Jul. 1959; C.P. Hoyt leg.; NHMUK. South Africa • 1♂; Dukuduku, between St Lucia and Matubatuba; 7-8 Apr. 1960; B. Stuckenberg leg.; NMSA • 1♀; Empangeni; 15 Apr. 2003; P. Reavell leg.; NMSA • 1♀; False Bay Park Reserve, Mpophomeni Trail area; 2 Jan. 1996; J. and A. Londt leg.; NMSA • 1♂ 1♀; Hazyview, Kruger Park Lodge area; 1-7 Oct. 2010; J. and A. Londt leg.; NMSA • 1♀; Middlekop; 11 Mar. 1979; NMSA • 1♂; Mpenjati Nature Reserve; 20 Mar. 2010; J. and A. Londt leg.; NMSA • 2♂♂ 1♀; same collection data as for preceding; 22-23 Jan. 2011 • 1♂; San Lameer Resort area; 11 Feb. 2012; J. and A. Londt leg.; NMSA. Tanzania • 1♀; Amani; G. Pringle leg.; NHMUK • 1♂; Kibonoto, Kilimanjaro; 29 Apr. 1906; Sjöstedt leg.; MNB • 1♀; Kigoma; 1924; C.R. Steel leg.; NHMUK • 1♂; Lembene, Usambara; 7 Jun. 1916; T.J. Anderson leg.; NHMUK • 3♀♀; Morogoro; J.G. Halcrow leg.; NHMUK • 1♂ 1♀; Morogoro; 16 May 1925; A.H. Ritchie leg.; NHMUK • 1♂ 1♀; same collection data as for preceding; 18 May 1925 • 1♂ 2♀♀; same collection data as for preceding; 20 May 1925 • 2♂♂ 1♀; same collection data as for preceding; 28 Jun. 1925 • 1♀; same collection data as for preceding; 10 Jun. 1925 • 1♀; Morogoro, Uluguru; A.G. Wilkins leg.; NHMUK • 1♀; Moschi; 1904; Alluaud leg.; MHNH. Togo • 1♀; Dzogbégan; 3 Jul. 1997; G. Goergen leg.; IITA • 1♂; Dzogbégan; 24-25 Jan. 2016; K. Jordaens leg.; KMMA • 1♂ 1♀; Kloto; 5 May 1996; G. Goergen leg.; IITA • 1♀; same collection data as for preceding; 3 Jul. 1997 • 1♂; same collection data as for preceding; Mar. 2000 • 6♂♂ 1♀; same collection data as for preceding; Aug. 2003 • 13♂♂ 5♀♀; same collection data as for preceding; Jan. 2004 • 3♂♂; same collection data as for preceding; Feb. 2004 • 31♂♂ 3♀♀; same collection data as for preceding; Mar. 2004 • 5♂♂ 1♀; same collection data as for preceding; Apr. 2004 • 1♂ 1♀; same collection data as for preceding; Jun. 2004 • 2♂♂; same collection data as for preceding; Nov. 2004 • 1♂ 1♀; same collection data as for preceding; Jan. 2005 • 4♂♂; same collection data as for preceding; Feb. 2005 • 3♂♂; same collection data as for preceding; Mar. 2005 • 1♀; same collection data as for preceding; Sep. 2005 • 1♀; same collection data as for preceding; Nov. 2005 • 1♂; same collection data as for preceding; Dec. 2005 • 1♂; same collection data as for preceding; Feb. 2006 • 1♀; same collection data as for preceding; May 2007 • 1♂; same collection data as for preceding; Jun. 2007 • 2♂♂; same collection data as for preceding; Oct. 2007 • 1♂; same collection data as for preceding; Jan. 2008 • 1♀; same collection data as for preceding; Jul. 2008 • 1♂; same collection data as for preceding; Jul. 2015 • 1♀; Kloto Forest; Jun. 2004; G. Goergen leg.; IITA • 1♂; Kuma-Tokpli; 21-24 Jan. 2016; K. Jordaens leg.; KMMA • 1♂; Lokossa; Jun. 2006; G. Goergen leg.; IITA • 1♀; Obé, near Akloa; 22 Apr. 2008; A. Ssymank leg.; MZH. Uganda • 1♀; Ankole; H.B. Johnston leg.; NHMUK • 1♂; Southeast Ankole; 4-8 Oct. 1911; S.A. Neave leg.; NHMUK • 1♂; Western Ankole; 10-14 Oct. 1911; S.A. Neave leg.; NHMUK • 1♀; between Jinja and Busia or Mbwago, E. Busoga; 28 Jul.-1 Aug. 1911; S.A. Neave leg.; NHMUK • 2♀♀; between Lake Kioga SE shore and Kakindu; 22-23 Aug. 1911; S.A. Neave leg.; NHMUK • 1♂; Buamba Forest, Semliki Valley; 3-7 Nov. 1911; S.A. Neave leg.; NHMUK • 3♂♂ 4♀♀; Budongo Forest, nr Lake Albert; Apr. 1972; E. Babyetagara leg.; CNC • 1♂ 1♀; Busoga; 1903; D. Bruce leg.; NHMUK • 2♂♂; Busoga; Mar. 1906; A. Hodges leg.; NHMUK • 1♂; Entebbe; 12-20 Jan. 1912; S.A. Neave leg.; NHMUK • 3♀♀; Entebbe; 14 Nov. 1912; C.C. Gowdey leg.; NHMUK • 1♂ 1♀; same collection data as for preceding; 18-20 Nov. 1912 • 1♀; same collection data as for preceding; 3-4 Dec. 1912 • 1♂; same collection data as for preceding; 12-13 Dec. 1912 • 1♀; Entebbe; 1 Feb. 1972; H. Falke leg.; CNC • 1♂; same collection data as for preceding; 10 Jun. 1972 • 1♂; same collection data as for preceding; 23-31 Jan. 1973 • 3♂♂; same collection data as for preceding; 25-28 Jan. 1973 • 1♀; same collection data as for preceding; Mar. 1973 • 1♀; Entebbe; 1 May 1973; M.K. Paulus leg.; CNC • 1♂; nr Entebbe; 23-31 Jan. 1973; H. Falke leg.; CNC • 1♂ 1♀; same collection data as for preceding; 1-14 Feb. 1973 • 1♀; south side Lake George; 17-19 Oct. 1911; S.A. Neave leg.; NHMUK • 4♂♂; Ibanda; 23-28 Dec. 1972; H. Falke leg.; CNC • 1♀; Jinja; Sep. 1928; V.G.L. van Someren leg.; NHMUK • 1♀; near Kakindu, Nile banks; 24 Aug. 1911; S.A. Neave leg.; NHMUK • 3♂♂ 2♀♀; Kayonza Forest, Kigezi; Sep. 1972; H. Falke leg.; CNC • 1♂; Kigaloma, Katwe; 2 Feb. 1912; H.B. Owen leg.; NHMUK • 1♀; Kilembe, Ruwenzori Range; Dec. 1934-Jan. 1935; F.W. Edwards leg.; NHMUK • 1♀; Magyo, Buvuma Island, Lake Victoria; Mar. 1968; E. Vertriest leg.; KMMA • 1♀; Masaka, Katera Forest; Nov. 1955; V.G.L. van Someren leg.; NHMUK • 1♀; Mbale-Kimu Road; 15-17 Aug. 1911; S.A. Neave leg.; NHMUK • 1♀; Mt Kokanjero; 7-9 Aug. 1911; S.A. Neave leg.; NHMUK • 1♀; W. Nile; 13 Nov. 1958 NHMUK • 2♂♂; Nkokonjera; 23 Dec. 1910; C.C. Gowdey leg.; NHMUK • 1♂; Nyabushozi, Nyrbthozi County; 21 Jan. 1975; P. Mugabi leg.; CNC • 1♀; Salt Lake to Wawamba Country; 1894; G.F. Scott Elliot leg.; NHMUK • 1♂ 1♀; Siroko River; 12-14 Aug. 1911; S.A. Neave leg.; NHMUK • 4♂♂ Tero Forest, Southeast Buddu; 26-30 Sep. 1911; S.A. Neave leg.; NHMUK • 1♂; Unyoro, Kafu River Valley; 23-28 Dec. 1911; S.A. Neave leg.; NHMUK • 1♂; Wasa River; 11-14 Apr. 1914; R.E. McConnell leg.; NHMUK. Zimbabwe • 1♀; Bomponi; 6 May 1963; D. Cookson leg.; NMSA.

Description.

Body length: 10.3-15.9 mm. Wing length: 7.9-12.0 mm.

Male (Fig. 1 ). Head (Figs 12 , 13 ). Eye bare; holoptic, eye contiguity for distance equal to length of ocellar triangle, facets dorsally slightly larger, at most twice as large in diameter as ventral ones. Frons black to black-brown; largely subshiny, with greyish brown pollinosity in dorsal fifth and along eye margins; with very dispersed, short pale pile. Face black to black-brown; subshiny with pale brownish to greyish pollinosity, in parts more densely so, medial part and ventral margins largely devoid of pollinosity; in parts with dispersed long pale pile; facial tubercle strongly pronounced. Gena colour and pollinosity as ventral lateral margins of face, with short to long pale pile. Occiput black-brown, covered with dull grey pollinosity; with dispersed pale pile. Antennal segments black-brown, basoflagellomere apically slightly paler; arista pale to dark yellow.

Thorax (Fig. 52 ). Scutum subshiny black; grey pollinose, except on anterior part of scutellum, at transverse suture and along lateral margins where more densely orange-brown pollinose; with short pale pile, anteriorly and on notopleuron somewhat longer. Scutellum pale brownish to orange-brown, anterior margin narrowly darker; with short pale pile. Pleura ground colour black-brown, sparsely greyish pollinose; covered with dispersed long pale pile, anterior anepimeron sometimes partially black pilose; pilosity absent on meron, dorsomedial anepimeron, anterior part of katepisternum and anterior anepisternum. Scutellum apical margin weakly rounded, distinctly marginated, 2.5 times as wide as long.

Legs. Brown to black-brown; with short black pile, along posterior margin of pro- and mesofemora with longer pale pile. Metafemur (Fig. 80 ) distinctly thickened, with one distinct ventral swelling in apical fifth, with dense short stout setae on swelling; metatibia thickened and slightly curved, with pile along ventral margin not distinctly more dense.

Wing (Fig. 64 ). Faint yellowish brown tinge, towards posterior margin and apex more greyish; with distinct dark brown macula running from anterior margin and covering most of stigma, parts of cell r1 and r2+3, distal part of cell br and basal part of r4+5, decreasing in colour on anterior part of cell dm; distally of brown macula more hyaline patch. Calypters yellow-white, in medial part more pale brownish; with fringe of yellow-white pile. Cell r1 variable, usually closed with petiole at most equal to height of base of stigma but usually shorter, sometimes closed but petiole missing; vein R4+5 sinuate, usually not appendiculate, rarely with short appendix.

Abdomen (Fig. 91 ). Subshiny brown to black-brown, postabdomen more reddish brown; with greyish to greyish brown pollinosity, except for anteromedial macula on terga I and II, pair of sublateral maculae on terga I-III and posterior part of tergum IV where more shining; with short dark pile, except tergum I, anterior margin tergum II, and lateral margins of all terga where pale pile. Sterna orange-brown to black-brown, with short dispersed pale pile. Male genitalia as in Fig. 102 .

Female. As male except for the following character states: Eye (Figs 14 , 15 ) dichoptic, facets equal to subequal in size. Frons subshiny black to black-brown in ventral protruding part, dorsally with greyish to greyish brown pollinosity. Postabdominal terga often less reddish brown; subshiny maculae sometimes more extensive and more pronounced.

Distribution.

Angola, Benin, Burundi, Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Ethiopia, Gabon, Ghana, Guinea Bissau, Ivory Coast, Kenya, Liberia, Malawi, Mozambique, Nigeria, Republic of the Congo, Sierra Leone, South Africa, Tanzania, Togo, Uganda, Zimbabwe. ‘Java’ as type locality is probably an error.

Comments.

The orginal description of Protylocera dentipes Macquart mentioned the geographic origin of the type as "de Java". No label on the lectotype pin indicates a locality or region, and the locality is probably an error. The type specimen of Protylocera dentipes bears a lectotype label with designation by F.C. Thompson in 1977. The original description did not indicate the number of specimens examined by Macquart but only made reference to the fact that the description is based on male only and that material is housed in the ‘Muséum’ (referring to MNHN). No additional specimens could be traced in the MNHN collection. The lectotype designation (made in accordance with recommendation 73♀ of the ICZN (Thompson pers. comm.)) has not been published elsewhere.

The type of Plagiocera maculipennis Loew could not be traced. According to the original description ( Loew 1858) as well as the detailed redescription by Loew (1860), this is a male specimen from ‘Guinea’. One female specimen in MNB collection (Guinea, Westerm., additional label ‘843’) is indicated as type but this contradicts with the above, and thus appears not be part of the type series. This specimen was also mentioned by Karsch (1887). According to Smith and Vockeroth (1980) ‘Guinea’ refers to Guinea Bissau.

The taxon Protylocera livida Bezzi was described as a variety of Plagiocera aesacus Walker by Bezzi (1912), based on two specimens from Fernando Po [= Bioko Island], which were among a series of specimens identified as Plagiocera aesacus . He pointed out the few morphological differences and suggested that they could be just teneral specimens, which did not attain their full colours and wing markings. Smith and Vockeroth (1980) elevated Plagiocera livida to species rank. Examination of the type material, however, indicates that Bezzi was correct in considering these as teneral specimens and that there is no morphological evidence for considering Plagiocera livida as a distinct species, different from Plagiocera dentipes . We, therefore, synonymize Plagiocera livida with Plagiocera dentipes .

Figures

Figures 1, 2. Senaspis species, habitus, lateral view 1 S. dentipes (Macquart) (♂) 2 S. dibapha (Walker) (♂).

Figures 12 - 15. Head, frontal and lateral view S. dentipes (Macquart) (12, 13 ♂ 14, 15 ♀).

Figures 52 - 55. Senaspis species, thorax, dorsal view 52 S. dentipes (Macquart) (♂) 53 S. dibapha (Walker) (♂) 54, 55 S. elliotii Austen (♂).

Figures 64 - 67. Senaspis species, right wing 64 S. dentipes (Macquart) (♂) 65 S. dibapha (Walker) (♂) 66, 67 S. dibapha (Walker) close-up calypters.

Figures 80 - 85. Senaspis species, hind leg, anterior view 80 S. dentipes (Macquart) (♂) 81 S. dibapha (Walker) (♂) 82 S. elliotii Austen (♂) 83 S. flaviceps Macquart (♂) 84 S. flaviceps Macquart (♀) 85 S. haemorrhoa (Gerstaecker) (♂).

Figures 91 - 96. Senaspis species, abdomen, dorsal view 91 S. dentipes (Macquart) (♂) 92 S. dibapha (Walker) (♂) 93 S. elliotii Austen (♂) 94 S. flaviceps Macquart (♂) 95 S. haemorrhoa (Gerstaecker) (♂) 96 S. melanthysana (Speiser) (♂).

Figures 102 - 113. Senaspis species, male terminalia 102 - 108, 110, 112 dorsal view 109, 111, 113 lateral view 102 S. dentipes (Macquart) 103 S. dibapha (Walker) 104 S. elliotii Austen 105 S. flaviceps Macquart 106 S. haemorrhoa (Gerstaecker) 107 S. nigrita (Bigot) 108, 109 S. melanthysana (Speiser) 110, 111 S. umbrifera (Walker) (holotype) 112, 113 S. nr umbrifera (Walker).