Antipodosis australis, Jaschhof, 2016

Jaschhof, Mathias, 2016, New species of Monepidosis Mamaev, 1966 and Antipodosis gen. nov., a closely related genus from New Zealand (Diptera, Cecidomyiidae), European Journal of Taxonomy 192, pp. 1-24 : 5-7

Antipodosis australis gen. et sp. nov.

urn:lsid:zoobank.org:act:45076E74-E261-4A53-804F-8264851F5133

Fig. 2 A–B

Diagnosis

A combination of genitalic characters is specific to A. australis sp. nov., as follows ( Fig. 2 A–B). Of the gonocoxites ( Fig. 2A ), the processes are small but prominent (↓); the ventral emargination, whose shape is reminiscent of an inverted bell, has a broad glabrous rim that almost reaches the anterior gonocoxal margin (↓), and the apodemes are merged into a large subrectangular plate (↓). The ejaculatory apodeme has a distinct subapical extension (↓, Fig. 2B ).

Differential diagnosis

Antipodosis australis gen. et sp. nov. resembles A. pureora gen. et sp. nov. in having elongate, slender gonostyli; a long, plate-like gonocoxal apodeme; two pairs of unmodified parameral tusks, and a singlepointed, straight ejaculatory apodeme. Characters to differentiate between the two species are provided by the gonocoxal emargination, which in A. australis gen. et sp. nov. is bell-shaped ( Fig. 2A ) and in A. pureora gen. et sp. nov. U-shaped ( Fig. 5A ), and by the ejaculatory apodeme, whose base is flattened in A. australis gen. et sp. nov. (not visible in the ventral view depicted in Fig. 2B ) and widened in A. pureora gen. et sp. nov. ( Fig. 5B ).

Etymology

The name australis is a Latin adjective meaning ‘southern’, which refers to the distribution of this species in the two southern main islands of New Zealand.

Type material

Holotype

NEW ZEALAND: Ƌ, South Island , Buller , Lake Rotoroa, mixed podocarp / southern beech forest, 450 m, 15 Jan. 2001, Malaise trap “15”, Department of Conservation St. Arnaud ( NZAC).

Paratype

NEW ZEALAND: 1 Ƌ, Stewart Island, Christmas Village Hut, 46.74° S, 167.97° E, 18 Jan. 2000, Malaise trap, R.K. Didham ( NZAC).

Other characters

BODY LENGTH. 2.0 mm.

HEAD. Eye bridge 5–6 ommatidia long dorsally. Circumfila on flagellomeres 1 to 13–14. Neck of fourth flagellomere 1.8–1.9 times longer than node. Palpus as long as height of head.

WING. Length/width 3.0–3.1.

LEGS. Empodia rudimentary.

TERMINALIA ( Fig. 2 A–B). Ninth tergite subtrapezoid, medially unsclerotized, asetose. Gonostylus strongly tapered towards apex, pectinate tooth narrow ( Fig. 2A ). Gonocoxites with small protuberances on ventroposterior margin; apodemes protrude clearly beyond ventroanterior margin ( Fig. 2A ). Ventral parameral tusks bent dorsolaterally, dorsal tusks bent dorsally ( Fig. 2B ).

Figures

Fig. 2. Antipodosis australis gen. et sp. nov., Ƌ, holotype. A. Genitalia, ventral. B. Parameres and ejaculatory apodeme, ventral. Scale lines: 0.05 mm.

Fig. 5. Antipodosis pureora gen. et sp. nov., Ƌ, holotype. A. Genitalia, ventral. B. Parameres and ejaculatory apodeme, ventral. Scale lines: 0.05 mm.