Atholus philippinensis ( Marseul, 1854 )

Cruz, Ian Niel dela, Hartini, Sri, Dwibadra, Dhian & Ôhara, Masahiro, 2024, Contribution to the Knowledge of the Genus Atholus (Coleoptera: Histeridae: Histerinae: Histerini) from the Indonesian Archipelago, Species Diversity 29 (1), pp. 119-139 : 131

Atholus philippinensis ( Marseul, 1854)

( Figs 1B , 11B )

Hister philippinensis Marseul, 1854: 547 (type locality: Philippines).

Hister philippensis [sic]: Gemminger and Harold 1868: 771.

Hister sectator Lewis, 1901: 375 [synonymized by Bickhardt (1917: 194)].

Atholus philippinensis : Lewis 1906: 402 (listed); Lewis 1915: 55; Mazur 1984: 215; Mazur 1997: 132; Ôhara 1999: 32–36 (Taiwan); Mazur 2011: 106 ( Myanmar, Vietnam, Indonesia: Sumatra, Borneo, Java, India: Meghalaya, China: Hainan; catalogued); dela Cruz and Ôhara 2023a: 1–26 ( Philippines).

Atholus sectator : Lewis 1906: 402 (listed).

Hister (Atholus) philippinensis : Bickhardt 1910: 54 (catalogued); Bickhardt 1913a: 173 (Hoozan, Taihorin); Bickhardt 1917: 194 (catalogued); Desbordes 1917: 322 (keyed); Miwa 1931: 57 (Hoozan, Taihorin); Kamiya and Takagi 1938: 31.

Specimens examined. Thirty-nine specimens of undetermined sex. Sumatra Island : S . Sumatra , 600 m, S . W. Lamponge , Mt . Tanggamoes, Giesting , December 1934, Lieftinck and Toxopeus leg . (1 ex. MZB) . Java Island : Holand Indien ( Atholus philippinensis Mars. Det. H. Desbordes, 1924 ), unknown collector (1 ex. MZB); Tjilatj . [= Tjilatjap ], August 1925, unknown collector (4 exs MZB); Tjilatjap, Java, 10 November 1925, Drescher leg . (2 exs MZB); W. Java , Bandoeng, 17 January 1940, alt . 750 m, J . Olthof leg . (1 ex. MZB); Java, Preanger, N .O.I., Radjamandala, G. Pantjankan , 30 August 1936, alt . 400 m, F . C . Drescher leg . (1 ex. MZB); W . Java , Lembang, 18 April 1940, alt . 1200 m, J . Olthof leg (1 ex. MZB) . Borneo Island : West Kalimantan, Mt . Bawang, 250– 300 m, October 1990, unknown collector (10 exs SEHU, 11 exs MZB); same locality, August 1990, unknown collector, (3 exs SEHU, 3 exs MZB); same locality, April 1991 [IC-22- in06], unknown collector (1 ex. SEHU) .

Diagnosis. Atholus philippinensis can be recognized for its long and almost complete external subhumeral striae.

Distribution. India: Meghalaya, Myanmar, Vietnam, China: Hainan, Taiwan, Philippines, Indonesia: Greater Sunda (Sumatra, Java and Borneo Islands) ( Fig. 11B ).

Remark. Among the Indonesian Atholus , this species is by far the largest in size.

Figures

Fig. 1. Some Indonesian Atholus, dorsal habitus. A, Atholus bifrons (Marseul, 1854), IC-22-in01; B, A. philippinensis (Marseul, 1854), IC- 22-in06; C, A. coelestis (Marseul, 1857), IC-22-in02; D, A. myrmidon (Marseul, 1861), IC-22-in05; E, A. famulus (Lewis, 1892), IC-22-in03; F, A. singalanus (Marseul, 1880), IC-22-in07; G, A. torquatus (Marseul, 1854), IC-22-in08. Scale bar: 1.00mm.

Fig. 11. Map showing the occurrence of some species of Atholus from the Indonesian archipelago and its extra-limital distribution in the Oriental Region. A, Atholus coelestis; B, A. philippinesis; C, A. torquatus; D, A. bifrons.