Raskyella peckii var. peckii Grambast & Grambast, 1954

Trabelsi, Khaled, Sames, Benjamin, Wagreich, Michael, K ́ azm ́, Miklos, , Andrea Mindszen & Mart ́ ın-Clo, d Carles, 2021, A new diverse charophyte flora and biozonation of the Eocene bauxite cover-sequence at G ant (V ertes Hills, Hungary), Journal of Systematic Palaeontology 19 (7), pp. 541-563 : 544-546

Raskyella peckii var. peckii Grambast & Grambast, 1954

( Fig. 3A–H)

1954 Raskyella pecki sp. nov. Grambast & Grambast: 670, text-fig. 1a–c.

1957 Raskyella pecki Grambast : 358, pl. 5, figs 7–9.

1958 Raskyella pecki Grambast : 190, fig 87a–c; 191, text-fig. 88.

1959 Raskyella pecki Horn af Rantzien: pl. 19, figs 7–13.

1971 Raskyella peckii subsp. ganesensis Soulíe-M¨arsche : pl. 2, 1–5.

1981 Raskyella pecki Anadon & Feist : pl. 1, figs 1, 2; pl. 2, figs 3, 4.

1986 Raskyella pecki Riveline : pl. 37, figs 7–9.

1999a Raskyella pecki Mart́ın-Closas et al.: 11, pl. 6, figs 1–3.

Material. Up to 65 gyrogonites in sample G-2.4, and dozens in samples G-2.2 and G-2.3. Collection numbers of figured specimens: HNHM-PBO 1501–1508.

Description. Gyrogonites are ovoidal to ellipsoidal in shape, spherical to sub-prolate ( ISI 100–120) and of large size, 800–1050 M m in height and 750–1050 M m in width, showing laterally 7–10 (usually nine) convolutions ( Fig. 4). Spiral cells often flat ( Fig. 3A) to slightly convex ( Fig. 3C), or concave ( Fig. 3B), but regularly without any kind of ornamentation. Apex broadly rounded, truncated and flattened, with the spiral cells abruptly discontinuing and ending acutely in the apical periphery to be replaced by the development of five opercular cells, each obliquely disposed at the end of a spiral cell ( Fig. 3F). Germinated specimens ( Fig. 3G) show a rose-shaped apical opening. Internal casts of these gyrogonites were also found ( Fig. 3E).

Remarks. The contemporaneous unornamented gyrogonites of Raskyella peckii subsp. ganesensis Soulí e-M ¨arsche, 1971 from the Aquitaine Basin ( France) appear to represent a gyrogonite population of relatively smaller size within R. peckii var. peckii and both are here considered synonymous. However, supplementary research on the type material is needed to verify this synonymy, since the morphotype ganesesis is only known from the type locality.

Additionally, the subspecies Raskyella peckii subsp. meridionale Grambast, 1960 , is kept within the rank of subspecies due to its palaeogeographical restriction. The extremely large gyrogonites of this subspecies are limited to the southern-most biogeographical range of Raskyella peckii , i.e. Algeria ( Grambast 1960; Mebrouk et al. 1997) and the Betic Domain of the Balearic Islands (Mart́ın-Closas & Ramos 2005).

Distribution. This is the first record of R. peckii var. peckii in Hungary and central Europe. This variety is widely distributed in the Lutetian and lower Bartonian of southern Europe, mainly in France ( Grambast & Grambast 1954; Grambast 1958; Soulíe-M¨arsche 1971, 1974; Riveline 1984, 1986) and Spain (Anadon & Feist 1981; Anadon et al. 1992; Mart́ın-Closas et al. 1999a; Mart́ın-Closas & Ramos 2005). The total range of this variety (early Lutetian — late Bartonian) has been characterized in the Eastern Ebro Basin (Catalonia) by Mart́ın-Closas et al. (1999a) based on correlations with larger foraminifera (mainly Nummulites ). Furthermore, R. peckii var. peckii has been also reported from North Africa, i.e. in the lower Eocene of Algeria ( Gevin et al. 1974; Vianey-Liaud et al. 1994; Mebrouk et al. 1997), in the late lower Eocene–early middle Eocene of Tunisia (Abdeljaoued et al. 1984) and in the Lutetian of Libya ( Megerisi & Mamgain 1980).