Ancylomenes tosaensis ( Kubo, 1951 ) Okuno & Bruce, 2010

Okuno, J. & Bruce, A. J., 2010, Designation of Ancylomenes gen. nov., for the ‘ Periclimenes aesopius species group’ (Crustacea: Decapoda: Palaemonidae), with the description of a new species and a checklist of congeneric species *, Zootaxa 2372 (1), pp. 85-105 : 101

Ancylomenes tosaensis ( Kubo, 1951) comb. nov. ( Fig. 6C )

Restricted synonymy:

Periclimenes (Ancylocaris) tosaensis Kubo, 1951: 268–271 , figs. 7, 8.

Periclimenes (Harpilius) tosaensis . — Bruce 1966: 15–21, figs. 1, 2, 3a, 4a, b.

Periclimenes tosaensis . — Hayashi 1986: 100, 101, 261, unnumbered fig. — Bruce 2008b: 15–16 , figs. 9, 14D. Periclimenes sp. 4 . — Minemizu 2000: 53, unnumbered fig.

Material examined. CMNH-ZC 01779 , 1 male, CL 3.4 mm, Ose-zaki , Numazu , Suruga Bay, Honshu, Japan, 31 m , SCUBA, in association with Dofleinia armata , 20 November 2003, leg . T. Yanagisawa ; CMNH-ZC 01830 , 1 female CL 3.5 mm , CMNH-ZC 01831 , 1 male, CL 3.0 mm, Masaki , Miho Peninsula , Shimizu , Suruga Bay, Honshu, Japan, 21 m , SCUBA, in association with Dofleinia armata , 4 November 2003, leg. J. Okuno.

Type data. The male holotype is now placed at the Tokyo University of Marine Science and Technology (former Tokyo University of Fisheries ) .

Distribution. Type locality: Off Usa, Takaoka, Tosa Bay, Shikoku, Japan. Also known from Seychelles, South China Sea, East China Sea, Philippines, northwest Australia, and Fiji ( Li & Bruce 2006; Bruce 2008b).

Bathymetric range. Moderately deep waters at the depths of 21–100 m.

Host. The present paper records the host of A. tosaensis comb. nov. for the first time. The specimens from Suruga Bay were associated with the actiniarian, Dofleinia armata .

Remarks. Although Hayashi (1986) and Bruce (2008b) provided the coloration of Periclimenes tosaensis based on the dead fresh specimen, its detailed colour pattern in life has been unclear. We examined the specimens to be considered same species reported as Periclimenes sp. 4 by Minemizu (2000), and identified them with Ancylomenes tosaensis comb. nov. This result indicates that this species is characterized by numerous fine pale yellow spots on the carapace and abdomen, and third abdominal tergum with a conspicuous red patch just anterior to a white dorsal patch (see Fig. 6C ). Previous identification of the individuals from Seychelles, Philippines and Fiji are possibly incorrect (Bruce in prep.).

Figures

FIGURE 6. A, B, Ancylomenes luteomaculatus sp. nov. Holotype, male, CL 3.6 mm, CMNH-ZC 02200. A, fresh specimen, lateral view (photo J. Okuno); B, same, dorsal view (photo J. Okuno). C, Ancylomenes tosaensis (Kubo, 1951) comb. nov. Living individual in situ, in association with Dofleinia armata Wassilieff, 1908 at Ose-zaki, Suruga Bay, Japan, 31 m, 20 November 2003 (photo T. Yanagisawa).