Enicospilus formosensis (Uchida, 1928)*

Shimizu, So, 2020, The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species, Deutsche Entomologische Zeitschrift 67 (1), pp. 69-126 : 69

Enicospilus formosensis (Uchida, 1928)* Figs 2E , 10

Henicospilus formosensis Uchida 1928: 223; holotype ♀, Taiwan, SEHU, examined.

Enicospilus saepis Chiu 1954: 77; holotype ♀, Japan, TARI, examined; synonymised by Gauld and Mitchell (1981: 424).

Material examined.

2♀♀ 2♂♂ and 1 unsexed: Nepal (1♂), Brunei (1♂), India (1 unsexed), Japan (1♀), Taiwan (1♀) .

Type series: holotype of Henicospilus formosensis Uchida , 1928, ♀, Baibara, Taiwan, 15.VI.1926, Y. Saito & Kikuchi leg. (SEHU) ; holotype of Enicospilus saepis Chiu , 1954, ♀, Nara, Honshû, Japan, 17.VIII.1918, J. Sonan leg. (TARI) .

Non-type series: 1♂, mixed forest (1,550 m), Godaveri , Nepal, 6.V.1984, M.G. Allen leg. (LT) (Figs 2E , 10 ) ; 1♂, Ulu Temburong (1,000 m), Brunei, II.1980, M.G. Allen leg.; 1 unsexed, NW Himalaya, Dalhousie, India, 8.VII.1965, Tikar leg. (all NHMUK) .

Distribution.

Eastern Palaearctic and Oriental regions ( Yu et al. 2016). Newly recorded from Nepal.

Diagnosis.

Head (Fig. 10B-D ): GOI = 2.2-2.4; lower face 0.8-0.9 × as wide as high; clypeus moderately convex in profile, its lower margin subacute to blunt; mandible weakly twisted by 10-20°, moderately long, evenly tapered, its outer surface without a diagonal structure; upper mandibular tooth 1.2-1.4 × as long as lower one; posterior ocellus close to eye; antenna with 66-69 flagellomeres and 20th flagellomere 2.1-2.2 × as long as wide.

Mesosoma (Fig. 10E ): mesopleuron moderately punctate; scutellum with lateral longitudinal carinae reaching anterior 0.8 or more and weakly convergent posteriorly so that subquadrate (Fig. 2E ); metapleuron punctate with isolated striae; propodeum declivous in profile, its posterior area coarsely irregularly wrinkled, sometimes with posterior transverse carina laterally, outer margin of propodeal spiracle joining pleural carina by a ridge or not.

Wings (Fig. 10F ): fore wing with AI = 0.2-0.6, CI = 0.2-0.9, ICI = 0.5-0.6, SDI = 1.1-1.3; fore wing vein 1m-cu&M weakly sinuous, 2r&RS almost straight; fenestra and sclerites of discosubmarginal cell of fore wing as in Figure 10F ; fenestra of fore wing not very long and its anterodistal corner distinctly separated from proximal end of vein RS; proximal sclerite triangular, confluent with distal one, strongly pigmented; central sclerite moderately to strongly pigmented and sclerotised, linear and parallel to distal margin of fenestra, positioned in distal part of fenestra; distal sclerite present proximally and vestigial to absent distally; proximal corner of marginal cell of fore wing uniformly setose; vein 1cu-a subinterstitial to antefurcal to M&RS by less than 0.2 × 1cu-a length.

Colour (Fig. 10 ): body including interocellar area testaceous; wings weakly infumate.

Differential diagnosis.

Enicospilus formosensis is a distinctive species and can easily be distinguished by many characters, such as the wide face (Fig. 10B ), shape of the central sclerite (Fig. 10F ), more or less subquadrate scutellum (Fig. 2E ), as listed in the diagnosis.

Figures

Figure 2. Diagnostic characters for some species of Nepalese Enicospilus. A-D. Outer surface of mandible: A. E. nepalensis Shimizu sp. nov., B. E. tangi Shimizu sp. nov., C. E. tripartitus, D. E. yonezawanus; E-H. Scutellum, in dorsal view: E. E. formosensis, F. E. kakanicus Shimizu sp. nov., G. E. phulchokiensis Shimizu sp. nov., H. E. tangi Shimizu sp. nov.; I, J. Hind tarsal claw: I. E. biharensis, J. E. nikami Shimizu sp. nov.

Figure 10. Enicospilus formosensis (Uchida, 1928), ♂. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing.