Tamaulipeca candoshi, Bordera, Santiago & Saeaeksjaervi, Ilari E., 2012

Bordera, Santiago & Saeaeksjaervi, Ilari E., 2012, Western Amazonian Ateleutina (Hymenoptera, Ichneumonidae, Cryptinae), Journal of Hymenoptera Research 29, pp. 83-118: 106-107

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Tamaulipeca candoshi

sp. n.

Tamaulipeca candoshi  ZBK  sp. n. Figs 9A, 9E, 9H

Material examined.

Holotype: ECUADOR, ♀, Dept. of Orellana, Onkone Gare, 0°39'25.7"S, 76°27'10.8"W, Canopy fog., 216.3m, 29.VI.1994, T. L. Erwin et al. leg., Lot#757 (USNM).


Female. Body length (without ovipositor) 5.0 mm, head length × width 0.8 × 1.2 mm, mesosoma length × width (mesoscutum) 2.4 × 0.7 mm, length of ovipositor sheath 1.8 mm, fore wing length 4.1 mm, antenna broken.

Head. Transverse, 0.40 times as long as wide, strongly narrowed behind eyes, gena at same level as hind rim of eye (dorsal view). Antenna broken; its segments 1, 4, 7 and 12 (and most of following segments) about 7.40, 5.00, 2.00 and 1.10 times as long as wide, respectively; basal flagellar segments 1 to 6 compressed or slightly cylindrical and flagellomere 7 and following depressed (flattened ventrally and this flattened area covered with short sensory setae); maximum width of flagellum about twice minimum width of first flagellomere; flagellum strongly tapered towards apex (Fig. 9A). Clypeus convex, almost triangular, with apex produced into acute point, about once as long as distance between c1ypeal foveae (Fig. 9E). Mandible short, about twice as long as width at middle, lower tooth slightly longer than upper tooth. Malar space about 0.70 times as long as basal width of mandible (Fig. 9E). Occipital carina absent dorsally, ventrally joining hypostomal carina just before mandible. Ocellar-ocular distance and distance between hind ocelli about once maximum diameter of lateral ocellus. Face coriaceous, matt and with moderately long silvery hairs (Fig. 9E). Frons matt, coriaceous. Vertex slightly coriaceous, with very sparse short hairs, lower vertex and occiput centrally concave; gena slightly coriaceous, occiput and gena with scattered silvery hairs.

Mesosoma. Upper margin of pronotum not swollen. Lateral part of collar without carina. Pronotum coriaceous, laterally slightly longitudinally strigose and with sparse silvery hairs. Mesoscumum coriaceous, slightly shiny, not strigose in the area between posterior end of notauli. Notauli reaching tegula level, convergent, strongly impressed and without clear transverse wrinkles along its length. Prescutelar groove with short longitudinal wrinkles. Scutellum weakly coriaceous, with lateral carinae present only at anterior end of the scutellum. Mesopleurum coriaceous, with very dense punctures and white long hairs. Mesopleural fovea present far in front of mesopleural suture. Sternaulus inconspicuous. Metapleurum and propodeum densely punctate or rugose-punctate on a coriaceous background, with dense whitish hairs. Posterior transverse carina of mesosternum complete (not interrupted infront of each middle coxa). Propodeum long, its spiracle round and very small. Anterior transverse carina absent. Posterior transverse carina present in the centre and in area posteroexterna, absent laterally, very slightly undulating along its length. Propodeum with short transverse carinae just before mid-length of the propodeum. Median longitudinal carinae of propodeum absent; lateral longitudinal carina absent anterior to posterior transverse carina, present behind it. Pleural carina strong. Area petiolaris confluent with areas posteroexterna. Vein 2rs-m short, about 0.40 length of M. Vein 2m-cu almost vertical and with one short bulla. M+Cu of hind wing strongly arched. Hind wing with distal abscissa of Cu1 strongly pigmented. Vein cu-a about same length as abscissa of Cu between M and Cu1 (Fig. 9H). Distad abscissa of 1A of hind wing short. Base of hind coxa of moderate depth, with a short transverse shallow groove next to its attachment. Tibiae and tarsi with sparse and stout bristles, bristles stronger on hind leg.

Metasoma. First tergite smooth, polished and about 1.70 times as long as maximum width; its maximum width at apex about 3.00 times minimum width (at base). Median dorsal and dorsolateral carinae of first tergite absent, ventrolateral carinae more or less distinct posteriorly. Spiracle at mid length of first tergite. Tergites 2-3 smooth and polished, laterally with short, sparse hairs; tergites 4-8 rather densely pubescent. Thyridium present as short impressed and granulate furrow. Ovipositor sheath 0.74 times as long as hind tibia, truncate at apex. Ovipositor moderately slender and slightly down curved (Fig. 9A), its tip elongate lanceolate, nodus weak, its lower valve without oblique ridges at apex.

Coloration (Fig. 9A). Head dark brownish with mandibles (except of brown teeth), palpi, clypeus mostly white or creamy white; orbits broadly whitish and gena with whitish spots. Antenna dark brown with scape yellowish and broad white band on flagellar segments 7-18. Mesosoma reddish with pronotum, mesoscutum and scutellum brown; pronotum with anterior edge broadly white, mesoscutum with 2 broad V-shaped marks and scutellum with white V-shaped mark. Legs mostly light brown with darker lighter coloration mostly in trochanters and hind tarsi. Metasoma dark brown with broad whitish bands on hind edges of tergites 1-2 and with a very narrow apical whitish band on tergites 4-6; tergite 7 whitish. Ovipositor sheath dark brown, ovipositor yellowish.

Male. Unknown.

Taxonomic discussion.

This speciesdiffers from all other species of the genus by the following characters in combination: mesoscutum and scutellum brown, with white V-shaped marks; metasoma dark brown with broad whitish bands on hind edges of tergites 1-2 and with a very narrow apical whitish band on tergites 4-6, tergite 7 whitish; and hind wing vein cu-a of about same length as abscissa of Cu between M and Cu1.


This species is dedicated to the Ecuadorian Candoshi people, who live in the Department of Orellana, Ecuador.


Only one specimen is known, collected in June.




The holotype was collected by canopy fogging in Onkone Gare, Department of Orellana, Ecuador, in the same habitat as Ateleute huaorani  .