Valdiviglypta, Broad & Sääksjärvi & Veijalainen & Notton, 2011

Broad, Gavin R., Sääksjärvi, Ilari E., Veijalainen, Anu & Notton, David G., 2011, Three new genera of Banchinae (Hymenoptera: Ichneumonidae) from Central and South America, Journal of Natural History 45 (21 - 22), pp. 1311-1329 : 1324-1326

Genus Valdiviglypta gen. n. Broad, Notton, Sääksjärvi & Veijalainen

( Figures 9 , 10 )

Type species: Valdiviglypta nimbus Broad, Notton, Sääksjärvi & Veijalainen. Monotypic.

Diagnosis

Valdiviglypta is a striking genus, with its large size, high and rather complete carination on the propodeum, strongly sculptured first metasomal tergite and a sternaulus-like groove that originates on the mesosternum.

Description

Large (body length c. 17 mm, excluding ovipositor, wing length 16 mm), mostly black with strongly infuscate wings. Clypeus flattened, slightly concave, but convex basally, 1.8 times as wide as high, lower margin rounded, with some long hairs medially and basally. Malar space 0.6 times basal width of mandible. Mandible wide basally, strongly, triangularly narrowed, with long, sparse hairs and lower tooth slightly shorter than upper. Face transverse and with prominent, rounded, central swelling. Margins of antennal sockets slightly raised. Low inter-antennal carina present, frons slightly convex, heavily sculptured, head depressed at margins of ocelli, with well-defined grooves adjacent hind ocelli. Temples strongly narrowed, eyes large. Ocellar–ocular distance 1.5 times maximum diameter of lateral ocellus. Occipital carina complete, laterally with slightly sinuous section, joining hypostomal carina distinctly above base of mandible. Antenna slim, white-banded sub-apically. Scape a little longer than wide, truncate about 45 ◦ from transverse. Flagellomeres with placoid sensilla evenly distributed. Pronotum short, with well-developed transverse groove. Epomia strong, straight. Dorsal, posterior corner of pronotum slightly twisted, in dorsal view rounded and not entirely concealing spiracular sclerite. Notauli distinct to two thirds length of mesoscutum, central lobe of mesoscutum protruding. Mesopleurum bulging out and with deep concavities around episternal scrobe (linking to mesopleural suture) and posterior half of subtegular region. Epicnemial carina present, ending distant from front edge of mesopleurum, just above level of lower corner of pronotum. Mesosternum with crenulate groove (ventrally displaced sternaulus?) originating at sternal angle of epicnemial carina, ending about mid-point of mesopleurum, closer to division of mesopleurum and mesosternum. Mesepisternal sulcus deep and strongly crenulate. Episternum bulging ventrally relative to coxal insertions. Lateral sections of posterior, transverse mesosternal carina (near hind coxa) high, carina otherwise absent. Metapleurum with submetapleural carina obtusely triangularly broadened anteriorly, as shallow lobe with strong crenulae. Propodeum steeply declivous, with most carinae present and strongly developed ( Figure 10A ). Area superomedia confluent with area basalis. Area petiolaris not marked. Central sections of longitudinal median carinae, postero-lateral corner of area dentipara and hind rim of propodeum particularly strongly developed. Propodeal spiracle oval. Legs slender, lacking spines or pegs. Tarsal claws slender and weakly curved, each claw with three or four very short teeth. Tarsomeres cylindrical. Hind tibial spurs short, about 0.2 times length of hind basitarsus. Fore wing with vein 3rs-m longer than 2rs-m, areolet petiolate. Vein 2m-cu with two narrow, widely spaced bullae. Vein cu-a distal to base of Rs&M by 0.4 times length of cu-a and strongly inclivous. Hind wing with Cu1 present, about equidistant between 1A and M. Tergite I, in lateral view, with spiracle positioned at anterior 0.45 and strongly protruding. Dorsal longitudinal carinae very strongly raised and close together, ending in two rounded protrusions at 0.8 of tergite length ( Figure 10B ). Tergite I with deeply excavate areas laterally and posterior to spiracle. Lateral carina present posteriorly, blunt but strong. Sclerotized part of sternite I not fused with tergite I and ending at about 0.4 of length of tergite. Tergite II about as long as wide, with raised central, roughly triangular area and broad, shallow depressions laterally, deepest anteriorly ( Figure 10B ). Anterior edge of tergite II with depressed, lateral areas. Laterotergites II–III narrow, turned under, laterotergite IV weakly separated from tergite, posterior laterotergites not separated. Subgenital plate large, triangular, but not quite extending to posterior level of apical tergite, with membranous, narrow triangular area postero-medially. Ovipositor slightly curved but basically straight, with dorsal, apical notch, 0.6 times length of fore wing.

Etymology

Valdiviglypta is named after the type locality, Valdivia, and the genus name Glypta , as we consider this genus to be best placed in the tribe Glyptini . The gender of the genus name is feminine.

Figures

Figure 9. Valdiviglypta nimbus holotype female, whole insect.

Figure 10. Valdiviglypta nimbus holotype female, (A) propodeum, dorsal, (B) propodeum and first to third tergites of metasoma.