Wormaldia clauseni Muñoz-Quesada and Holzenthal, 2008

Muñoz-Quesada, Fernando J. & Holzenthal, Ralph W., 2008, Revision of the Nearctic species of the caddisfly genus Wormaldia McLachlan (Trichoptera: Philopotamidae), Zootaxa 1838, pp. 1-75 : 22-25

Wormaldia clauseni Muñoz-Quesada and Holzenthal , new species

Figures 27–33, 129

According to the species group proposals of Ross (1956) and Armitage (1996), this new species would be a member of the W. anilla Group (Table 1).

This new species is similar to Wormaldia anilla ( Ross 1941a) and W. occidea ( Ross 1938a) , but differing from these two species in the shapes of sterna VII and IX, tergum X, and the inferior appendage. Sternum VII in W. clauseni has a broad and convex process posteromesally (less than 0.2 times the length of sternum VIII). Sternum VII in W. anilla lacks a process posteromesally, in W. occidea there is an elongate, broad, convexly subtriangular process posteromesally (longer than posteromesal process of W. clauseni ). Sternum IX in W. clauseni has a stout, elongate, truncate process posteromesally that arises preapically. Sternum IX in W. anilla has a slender, pointed process posteromesally that arises preapically; in W. occidea the process is smaller and truncate posteromesally and arises subapically. Tergum X in W. clauseni is nearly triangular, broad, concave laterobasally, bulged medially, and widely rounded apically. Tergum X in W. anilla and W. occidea is triangular, straight laterally, and slightly pointed apically. When viewed laterally, the apical segment of the inferior appendage in W. clauseni is thick and very weakly concave medially, and with the apex uniformly and widely rounded. The apical segment of the inferior appendage in W. anilla is slightly concave medially and widely rounded apically; in W. occidea it is straight medially, and very slightly truncately rounded apically.

Adult. Length of male forewing 6 mm (holotype). Head brown, with yellowish setae. Antenna long, slen- der, brown, with small, lighter setae. Maxillary palps yellowish, with lighter setae, fifth segment the longest, first segment the shortest, third longer than second and fourth, second and fourth subequal. Labial palps yellowish, with lighter setae, first and second segments sub-equal, third segment the longest. Dorsum of thorax brown. Legs yellowish, with small, lighter setae. Forewing yellowish, covered with fine, small, brown setae, with apical forks I, II, III, IV, and V present ( Fig. 32). Hind wing translucent, with very few fine, small, brown setae, with apical forks I, II, III, and V present ( Fig. 33).

Male genitalia ( Figs. 27–31). Sternum VII with broad, convex posteromesal process, less than 0.2 times length of sternum VIII. Tergum VIII straight or weakly concave posteriorly. Sternum VIII with small, broad, convex posteromesal process, about 0.1 times length of posteromesal process of sternum VII. Segment IX, when viewed dorsally, concave anteriorly; when viewed laterally, slender and nearly rectangular in appearance, convexly projected anteriorly, weakly sinuous posteriorly; when viewed ventrally, weakly concave anteriorly, nearly straight posteriorly with stout, elongate, truncate, mesal process arising preapically. Segment X, when viewed dorsally, nearly triangular, elongate, concave anterolaterally, bulged medially, narrowest and widely rounded posteriorly; when viewed laterally, stout, rounded apically. Superior appendages digitate; when viewed dorsally, parallel with segment X, stout, elongate, rounded apically; when viewed laterally, slightly shorter than segment X, slender, rounded apically. Inferior appendages two segmented; when viewed laterally, basal segment stout, rectangular, elongate, broadest medially, convex dorsally, relatively straight ventrally; apical segment stout, rectangularly elongate, nearly equal in length, somewhat narrower than basal segment, weakly concave medially, widened and rounded posteriorly; when viewed dorsally, apical segment stout, tubularly elongate, narrowest and rounded posteriorly, with elongate apicolateral patch of short, thin, black, spine-shaped setae on inner margin; when viewed ventrally, basal segments paired, united for about their anterior three-quarters, separated posteromesally by a shallow, broad, concave emargination, each basal segment thick, widest medially, with outer margin convexly curved, apical segment as when viewed dorsally. Phallus, when viewed laterally, pistol-shaped, widest basally, tapering from middle to apex, membranous api-

cally, very lightly sclerotized, with three visible, small, sclerotized, internal processes ( Fig. 31), surrounded by highly convulated membranes with a group of many minute, slender, pointed, internal sclerites.

Holotype: Male. CANADA: British Columbia: Agassiz [49º 14’ N, 121º 46’ W], 19.v.1927, H.H. Ross (in INHS, Type INHS # 16199 ).

PARATYPES: CANADA: same data as holotype except, 1 male (in UMSP, Type INHS # 16202 ); British Columbia: Cultus Lake , R. Ck. [49º 04’ N, 121º 58’ W], 12.v.1935, W.E. Ricker, 1 male (in INHS, Type INHS # 16200 ) ; same data except, 1 male (in NMNH, Type INHS # 16201 ) .

Etymology. This species is named in honor of Dr. Philip J. Clausen of the University of Minnesota Insect Collection (UMSP), who is an exceptional and happy curator and human being. The senior author is deeply thankful for his valuable collaboration, advice, and friendship. Muchas Gracias Felipe!

Distribution. Canada: British Colombia.