Cheumatopsyche agha Oláh & Johanson, 2008

Oláh, János, Johanson, Kjell Arne & Barnard, Peter C., 2008, Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera), Zootaxa 1738, pp. 1-171 : 105-107

Cheumatopsyche agha Oláh & Johanson , new species

Fig. 257–259

This species is similar to C. ayopa , new species but in C. agha the clefts separating the 2 harpagonal processes are shallower, about as long as one-third the length of the harpagones. A striking feature is also the setose process located on the dorsobasal part of segment X, which is reduced in C. ayopa . The apicomesal incision on segment X is narrower and shallower than in C. ayopa .

Male. Body and wings brown with brown pubescence; wing unicolorous without spots. Maxillary palp segment I shortest, segment II longer, segments III and IV equally long, segment V as long as sum of segments I–IV. Head dorsum brown with 7 similary coloured warts. Antennae about 1.5 times longer than forewings. Proepisternum without swollen setal wart. Protarsal claws asymmetrical, laterally flanked by setal bundle. Forewing length 8.8 mm, hind wing length 6.4 mm. Distance between forewing crossvein m-cu and cu about 1.5 times length of crossvein m-cu. Hind wing fork I present.

Male genitalia. Segment IX fused annularly; tergum and sternum short ( Fig. 257); anterior margins of segment IX symmetrically convex; apical lobe on posterolateral margins small, located immediately above base of coxopodites; spine row along posterior margin of segment IX complete ( Fig. 257); dorsal spines 2 times longer than other spines. Intersegmental step between segment IX and segment X sloping. Segment X long, curving dorsad from mid-length, distal half gradually narrowing ( Fig. 257); simple, nearly tongue-shaped in dorsal view ( Fig. 258). Apicoventral setal lobes completely fused to segment X, visible as area delineated by setae. Setaless mesocaudal lobe completely reduced ( Fig. 258). Apical incision on segment X minute. Large, setose process present on dorsobasal part of segment X, clearly visible in lateral view ( Fig. 257); in dorsal view apparently located on left side of segment X. Transverse sutures shifted to basalmost section of segment X ( Fig. 257). Longitudinal sutures running along ventral margin of segment X, merging with transverse sutures at basis of segment X. Setose areas forming pair of elevated warts in basal region of segment X. Coxopodites robust, basal halfs slender; strongly broadening from mid-length; slightly exceeding apex of segment X ( Fig. 257). Harpagones with bifid apices; cleft separating branches about as long as one-third harpagones length. Phallotheca ( Fig. 259) with broad phallobase, horizontal part slender; sclerotised endothecal process circular, curving ventrad (in cleared preparation of holotype, but not in paratype); phallotremal sclerites large, rounded, vertical in lateral view.

Holotype male: MADAGASCAR: Perinet , xii.1955 [R. Paulian] ( MNHN, in alcohol).

Paratype: MADAGASCAR: Andohahelo , 1500 m, i.1954 [R. Paulian] — 1 male ( OPC, in alcohol) .

Distribution: Madagascar.

Etymology: after the uniquely bifid and half-leaved harpago, half “agha” in Sanscrit.

Remark: The large, setose process on dorsobasal part of segment X, clearly visible in lateral view ( Fig. 257) is located on the left side of segment X, indicating that a right setose process has been reduced. The process may therefore represent the apicodorsal setose processes present in Hydropsyche , Orthopsyche , which have shifted basally.