Cheumatopsyche Wallengren, 1891

Oláh, János, Johanson, Kjell Arne & Barnard, Peter C., 2008, Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera), Zootaxa 1738, pp. 1-171 : 9-10

Cheumatopsyche Wallengren, 1891

Cheumatopsyche Wallengren, 1891: 142 . Type species (monobasic) Hydropsyche lepida Pictet, 1834: 207 .

Hydropsychodes Ulmer, 1905a: 34 . Type species (by designation of Mosely, 1939: 27) Ulmeria Navás, 1918: 15 . Type species (by original designation) Hydropsyche lepida Pictet, 1834 , (synonymised by Navás 1933: 97).

Ecnopsyche Banks, 1913: 179 . Type species (monobasic) Ecnopsyche reticulata Banks, 1913: 179 (synonymised by Mey 1997: 306).

The species in the genus resemble those in Potamyia , but in the hind wings the stems of M and Cu1 are neither parallel nor close together in Cheumatopsyche , and on posterior margin of segment IX in the genitalia a tongue-shaped apical lobe is well developed in several Cheumatopsyche species groups but never present in Potamyia . There are 5 to 9 setal warts on the head dorsum: a single fused pair of anteromesal (interantennal) setal warts; a pair of anterior (postantennal) setal warts; a pair of anterolateral setal warts, frequently divided into 2 separate warts or even diffused into a setal area with several individual small warts; and a pair of posterior (cephalic) setal warts dominating the entire dorsum. In the hind wings fork 1 is absent or present, and the median cell is open. The male protarsal claws are of various shapes, most often laterally flanked by a bundle of setae; some species, mainly from Madagascar, have perfectly symmetrical protarsal claws.

Male genitalia. The terminology applied to genitalic structures are given in Fig. 16–19. Segment IX is fused annularly, its apicolateral corners of tergum IX are slightly prominent; dorsocaudal spiny lobes are usually present as a pair on the dorsolateral corners; a single ventrocaudal lobe is present only in some species in various species groups. The dorsum of segment X is trilobed, with a setaless mesocaudal lobe forming a tongue-like plate on the apicodorsal margin, and a pair of apicoventral setal lobes. The apicoventral setal lobes are either developed into paired processes, as lobes, ridges, compact and diffused surfaces, or are completely fused with segment X. The setaless mesocaudal and setose apicoventral lobes encircle the various dorsal and lateral interlobular gaps, as seen in dorsal and lateral views. The reduction or fusion of these segment X lobes are a basis for delineating the Cheumatopsyche species groups. There are transverse and longitudinal sutures present on segment X, usually meeting anteriorly in a Y-shape suture pattern. Apicodorsal setose area or lobes are present in several hydropsychine genera ( Hydromanicus , Hydatopsyche , Hydatomanicus , Hydropsyche , Orthopsyche , Aoteapsyche , Caledopsyche ), but present only in a single Cheumatopsyche species group (the C. holzschuhi group). The gonocoxites are usually slender, and the various shapes of the harpagones are important diagnostically. The phallic apparatus is modified into a phallothecal tube with a thick base, and a simple termination with a pair of sclerotised endothecal processes, considered synonym with the endothecal papillae or valves of Schmid (1998) and outer lip of McFarlane (1976), covering the heavily sclerotised phallotremal sclerites and the highly reduced ventral membranous endothecal lobe.