Crassignatha shunani Y. Lin & S. Li, 2020

Li, Ya, Lin, Yucheng & Li, Shuqiang, 2020, A review of Crassignatha (Araneae, Symphytognathidae), ZooKeys 988, pp. 63-128 : 63

publication ID	https://dx.doi.org/10.3897/zookeys.988.56188
publication LSID	lsid:zoobank.org:pub:6E64D69B-DD73-4A7E-AE2B-3CD21247A5E3
persistent identifier	https://treatment.plazi.org/id/099E51E5-0B4A-4433-8BFA-995B8FB3E784
taxon LSID	lsid:zoobank.org:act:099E51E5-0B4A-4433-8BFA-995B8FB3E784
treatment provided by	ZooKeys by Pensoft
scientific name	Crassignatha shunani Y. Lin & S. Li
status	sp. nov.

Treatment

Crassignatha shunani Y. Lin & S. Li sp. nov. Figs 28 View Figure 28 , 29 View Figure 29 , 38 View Figure 38

Type material.

Holotype ♂ (NHMSU Ar 073) and paratypes 4♂ 49♀ (NHMSU Ar 074-126), China: Sichuan Province, Luzhou City, Gulin County, Jianzhu Township, Wenyi Village, Dahei Cave (28.06134°N, 105.58015°E; 852 m), 23.IV.2014, Y. Lin, H. Zhao and J. Wu leg.; 1♂ juvenile (NHMSU-HA046) and 1♀ (NHMSU-HA046) used for sequencing, GenBank: MT991994 and MT991993, same data as for preceding.

Diagnosis.

This species is similar to C. nantou sp. nov. in the form of the male palp and the vulva configuration but can be distinguished by having a sharp cymbial tooth and lacking a hooked process on the median apophysis (Fig. 29A View Figure 29 vs. Fig. 15A View Figure 15 ), and by the more widely separated spermathecae and lower inflection point of the copulatory ducts in the center of the vulva (Figs 29F View Figure 29 , 15G View Figure 15 ).

Description.

Male (holotype). Total length 0.68. Carapace 0.32 long, 0.28 wide, 0.32 high. Clypeus 0.12 high. Sternum 0.20 long, 0.20 wide. Abdomen 0.40 long, 0.40 wide, 0.48 high. Length of legs: I 1.02 (0.32, 0.12, 0.24, 0.16, 0.18); II 0.88 (0.24, 0.12, 0.20, 0.16, 0.16); III 0.66 (0.18, 0.10, 0.12, 0.10, 0.16); IV 0.76 (0.22, 0.10, 0.16, 0.12, 0.16).

Somatic characters (Fig. 28A-C View Figure 28 ). Coloration: prosoma brown. Legs pale brown. Abdomen grayish yellow dorsally, dark brownish laterally and ventrally, with weakly sclerotized dots dorsally. Prosoma: carapace sub-rounded, cephalic and thoracic area granular, thoracic center and clypeus smooth. Cephalic area strongly elevated. Clypeus concave. ALE protruded. PER distinctly recurved. Chelicerae bears short setae anteriorly. Labium subtriangular. Sternum heart shaped, surface textured, slightly swollen, truncated posteriorly. Legs: tibia II with two clasping spurs. Abdomen: nearly globular, posterolateral scutum weakly sclerotized. Spinnerets without circular plate.

Palp (Fig. 29A-C View Figure 29 ): tibia laminar, subequal to patella in length. Cymbium with some setae at distal margin, horn-shaped cymbial tooth near cymbial apex. Tegulum globose, swollen, surface rugose. Nearly rounded median apophysis prolaterally on bulb, with short, straight distal process. Embolic membrane arises beside median apophysis. Embolus long, spiral, basally and mesally wide, distally narrow and bent.

Female (one of paratypes). Total length 0.80. Carapace 0.36 long, 0.32 wide, 0.32 high. Clypeus 0.12 high. Sternum 0.24 long, 0.20 wide. Abdomen 0.56 long, 0.60 wide, 0.64 high. Length of legs: I 1.12 (0.38, 0.12, 0.26, 0.18, 0.18); II 0.94 (0.26, 0.12, 0.22, 0.14, 0.20); III 0.74 (0.18, 0.10, 0.16, 0.12, 0.18); IV 0.88 (0.26, 0.12, 0.18, 0.12, 0.20).

Somatic characters (Fig. 28D-F View Figure 28 ). Coloration: prosoma and legs as in male. Abdomen black, with sclerotized dots dorsally. Prosoma: carapace pear shaped, surface modification and arrangement of eyes as in male. Cephalic area lower than in male. Mouthparts and sternum as in male. Abdomen: globular. Spinnerets tiny, lacking circular plate.

Epigyne (Fig. 29D-G View Figure 29 ): epigynal area slightly sclerotized. Scape protruded, as wide as long. Copulatory openings located at terminus of scape. Internal structures faintly visible via translucent tegument. Spermathecae strongly sclerotized, separated by their diameter. Fertilization ducts originating posteromedially from spermathecae. Copulatory ducts long, connected to posterior margin of spermathecae, curved upward and inward to center of vulva, then turned sharply downward to copulatory openings. The base of copulatory ducts unfused.

Etymology.

The specific name is from the Chinese pinyin shŭ nán, referring to the collection locality of this new spider species from southern Sichuan; noun in apposition.

Distribution.

China (Sichuan) (Fig. 38 View Figure 38 ).