Cyclosomus Latreille, 1829

Kavanaugh, David H., Cueva-Dabkoski, Mollie & Liang, Hongbin, 2023, Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species., Proceedings of the California Academy of Sciences 67 (20), pp. 451-491 : 459

Genus Cyclosomus Latreille, 1829

Scolytus Fabricius, 1790:221 , in part [junior homonym of Scolytus Geoffroy, 1762 ] (type species Carabus lim-

batus Fabricius, 1801, designated by Latreille (1810:426)). Fabricius (1792:180); Bousquet (2012:89). Cyclosomus Latreille, 1829:394 (type species Carabus flexuosus Fabricius, 1775 , by monotypy).

Diagnosis. Adults of Cyclosomus can be distinguished from those of Tetragonoderus species in the study area by the following combination of character states: Body size larger, BL = 7.0 mm or more; pronotum ( Fig. 13A ) very broad, trapezoidal, almost as wide as elytra at humeral angles, distinctly widest at base; elytra broadly rounded apically, elytral color pattern as in Fig. 7A ; front tarsomeres 1 to 3 markedly expanded laterally ( Figs. 14A , 15A ) in both males and females.

Diversity: At present, only five species are recognized in the Asian fauna, and here we record one, a new species, from the study area.

Figures

FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm.

FIGURE 7. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.

FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm.

FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm.