Carpelimus (s. str.) indicus (Kraatz, 1859) Scheerpeltz, 1971

Gildenkov, Mikhail Yu., 2020, New data on the New Guinea fauna of Carpelimus Leach, 1819 (Coleoptera: Staphylinidae: Oxytelinae), Amurian Zoological Journal XII (3), pp. 369-377 : 371-373

Carpelimus (s. str.) indicus wamenensis Gildenkov, ssp. n.

http://zoobank.org/NomenclaturalActs/E8A- 3 BAF 6-67CB-4DE9-9538-10010CF23CFC

Material. Indonesia: Holotypus, ♂ “ Irian Jaya: Jayawi-jayajayag, Wamena, Beliem-Fluss 15.10.1993 leg. A. RIEDEL 1700 m ” ( SMNS) . Paratypus: 3♂, 4♀, 56 ex. “ Irian Jaya: Jayawi-jaya, Wamena, Beliem-Fluss 15.10.1993 leg. A. RIEDEL 1700 m ” ( SMNS; 1♂, 1♀, 5 ex. — cMG) .

Description (holotype). Length 3.0 mm (paratypes — from 2.9 to 3.2 mm; the length of the nominative subspecies is 2.6–3.0 mm). Colouration black-brown, antennae and legs brown, apices of tibiae and tarsi — yellow brown. Integument slightly shining, body with short, light-coloured hairs.

Head transverse, with a wide base, ratio of its length (from posterior margin of head to anterior margin of clypeus) to maximum width about 23:36 (in the nominative subspecies — 23:35). Neck constriction prominent. Eyes large, convex, occupying almost entire lateral side of head; temples almost indistinct, head widest across eyes (see Fig. 1 ). Head surface with delicate, fine and sparse punctation. Diameter of punctures is about 4.0 times as small as an eye facet. Distances between punctures are significantly more than their diameter, interspaces smooth, slightly shining. Antennae long, antennal segments 1–11 elongate. Last 3 segments more massive than the others and form a lose club.

Pronotum widest at about 2/3 of its length from base, then narrowed. Lateral margins straight from base, then smoothly rounded (see Fig. 1 ). Ratio of pronotum length to its maximum width about 34:46 (in the nominative subspecies — 31:43). Surface of pronotum with delicate, fine and rather rare punctation. Diameter of punctures about 4.0 times as small as an eye facet. Distances between punctures significantly more than their diameter, interspaces smooth, slightly shining. Pronotal disc with 2 pairs of prominent, symmetrical depressions and 1 unpaired oval depression along midline near apex. Base of pronotal disc with narrow, crescent-shaped depressions separated by a medial ridge. Central part of disc with oval depressions (see Fig. 1 ).

Ratio of length of elytra to their combined width about 53:57 (in the nominative subspecies — 48:53). Surface of pronotum with clear, large and dense punctation. Diameter of punctures is about equal to an eye facet. Distances between punctures are a little smaller than their diameter, interspaces smooth, shining.

Abdomen delicately shagreened.

Aedeagus of characteristic structure (see Fig. 2 ). Female. Sexual dimorphism is absent, female morphologically similar to male. Spermatheca has a characteristic structure (see Fig. 3 ) for representatives of the subgenus Carpelimus ( Gildenkov 2015) .

Differential diagnosis. It differs from the nominative subspecies with which the edeagus is identical in structure (see Fig. 2 ) by larger sizes (see above) and darker colour (see Figs. 1, 4 ). It is reliably distinguished by more rare punctation of the pronotum and significantly larger (about 1.5–2.5 times) punctation of the elytra (see Figs. 5, 6 ).

Distribution: Wamena (Jayawijaya regencies), largest town in Indonesian Papua's highlands, in the Baliem-river Valley.

Etymology. Named for its geographical distribution.

Remarks. For the nominative subspecies (see above), on the basis of the extensive material that we studied ( Gildenkov 2015; 2016), some variability is known in the width of the pronotum, puncture of the pronotum and elytra. However, so far the variability has been random. In the new subspecies, all 64 specimens have stable signs of colouration and body size, puncture of the pronotum and elytra. These characters differ significantly from those of the nominative subspecies, and this is the reason for the description of the new subspecies.

Figures

Figs. 1–4. Carpelimus indicus spp.: 1 — Carpelimus (s.str.) indicus wamenensis Gildenkov, ssp. n., holotype, male, habitus, dorsal view; 2 — aedeagus, ventral view; 3 — spermatheca, lateral view; 4 — Carpelimus (s.str.) indicus indicus (Kraatz, 1859), male (New Guinea, Irian Jaya Jayapura distr.), habitus, dorsal view. Scale bar: 2 — 0.25 mm.; 3 — 0.1 mm Рис. 1–4. Carpelimus indicus spp.: 1 — Carpelimus (s. str.) indicus wamenensis Gildenkov, ssp. n., гоΛотип, самец, габитус, виΑ сверху; 2 — эΑеагус, виΑ снизу; 3 — сперматека, виΑ сбоку; 4 — Carpelimus (s. str.) indicus indicus (Kraatz, 1859), самец (Новая Гвинея, про- винция Ириан-Àжая), габитус, виΑ сверху. Масштаб: 2 — 0,25 mm; 3 — 0,1 mm

Figs. 5–6. Carpelimus spp., punctation of the surface of the pronotum and elytra: 5 — C. indicus wamenensis Gildenkov, ssp. n., holotype; 6 — C. indicus indicus (Kraatz, 1859), male (New Guinea, Irian-Jaya, Jayapura distr.). Рис. 5–6. Carpelimus spp., пунктировка поверхности переΑнеспинки и наΑкрыΛий: 5 — C. indicus wamenensis Gildenkov, ssp. n., гоΛотип; 6 — C. indicus indicus (Kraatz, 1859), самец (Новая Гвинея, провинция Ириан-Àжая)