Lapsias cyrboides Simon, 1900

Maddison, Wayne P., 2019, A new lapsiine jumping spider from North America, with a review of Simon's Lapsias species (Araneae, Salticidae, Spartaeinae), ZooKeys 891, pp. 17-29 : 17

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Lapsias cyrboides Simon, 1900


Lapsias cyrboides Simon, 1900   Figs 30-40 View Figures 30–40

Type material.

In MNHN, 3 males, 4 females, 3 juveniles from Colonia Tovar, Aragua State, Venezuela, in a vial with label "20924 Laps. cyrboides   E.S., Tovar!" and a more recent label "det Szűts 0014". Galiano (1963) designated one male as a lectotype, in separate microvial with her label "Typus? M.E. Galiano II 1959". She mentions one female designated also as lectotype, but no female is separated and labelled as such. Because Galiano (incorrectly) designated two lectotypes, the name is not yet fixed to a single specimen. This ambiguity is resolved by designating her male lectotype as the only lectotype.


Simon (1900) described a male and female. However, as noted by Galiano (1963), there are two species of female among the four females in the type vial, similar in body but easily distinguished by the epigyne. Two of the females ( Figs 37-40 View Figures 30–40 ) have an anteriorly placed guide ( Fig. 37 View Figures 30–40 ), while the other two females ( Figs 48-51 View Figures 41–51 ) lack such a guide and instead show two wing-shaped atria extending laterally ( Fig. 48 View Figures 41–51 ). It is reasonable to assume that these two kinds of female belong to the two smaller-bodied Lapsias   at Colonia Tovar, L. cyrboides   and L. tovarensis   . Under L. cyrboides   Simon described the female kind with anterior guide ("Plaga genitalis...longior quam latior"), but he did not justify this choice nor even mention the second form of female. Galiano followed Simon’s choice of matching female. The two forms of female are approximately the same size and carapace shape and are too faded to supply distinctive markings by which to match to the males. Nonetheless, I tentatively support Simon’s and Galiano’s matching based on an expected correlation between the form of the female’s guide and that of the male’s RTA. An anterior guide is expected to be associated with an extraordinary RTA, for instance, as in Papuamyr omhifosa   Maddison & Szűts, 2019 ( Maddison and Szűts 2019). Males of L. ciliatus   and L. tovarensis   have small RTAs, suggesting a simple or small guide along the epigastric furrow, as is common in salticids, while the male of L. cyrboides   has an unusual dorsally projecting tibial apophysis, which predicts an unusually-placed female guide. Thus, the female with anterior guide is tentatively considered that of L. cyrboides   , and the female with wing-shaped atria is considered that of L. tovarensis   .