Macratria tongkonan, Telnov, 2023

Telnov, Dmitry, 2023, Fourth contribution to the Macratriinae LeConte, 1862 of the Papuan Region and Wallacea (Coleoptera: Anthicidae) with descriptions of sixteen new species, Zootaxa 5361 (1), pp. 1-52 : 49-51

Macratria tongkonan sp. nov.

urn:lsid:zoobank.org:act:68E9393C-68B4-4E31-8E61-B7BD98CF8D20

( Figs 138–147 )

Type material designated. Holotype ♂ NME: INDONESIA, Sulawesi, South Sulawesi Prov., Makale 6 km SSW, 3°08’S 119°49’E, 11.I.1998, 1700 m, disturbed lowland rainforest, day collecting [printed].

Paratypes 7♂ ♀ specimens. 3♂ ♀ NME & 2 DTC: same label as holotype ; 2♂ EMNH: INDONESIA SULAWESI 23.03. Gintu road 1°48’S / 120°22’E forest, sweep net bush leg. U.Roosileht [printed] .

Derivatio nominis. Named after Tongkonan, the traditional ancestral house of the Torajan people in South Sulawesi where the new species was found. Tongkonan have a peculiar boat-shaped, oversized saddleback roof. Noun in apposition.

Measurements, holotype male, total body length 4.4 mm; head length 0.9 mm, head width across compound eyes 0.7 mm, pronotal length 1 mm, maximum pronotal width 0.65 mm, elytral length 2.5 mm, combined maximum elytral width 1.05 mm. Selected male paratypes 4.5–4.7 mm, selected female paratype 4.3 mm long.

Description. Male ( Fig. 138 ). Dorsum and venter uniformly black. Mouthparts including maxillary palpi, legs and antennomeres 1–8 pale rufous to yellow, remaining antennomeres yellowish brown. Head moderately glossy dorsally and ventrally, elongate elliptical, distinctly narrows anteriad and towards base. Frons narrow, about half to slightly more as wide as dorsal eye length. Compound eye large, strongly protruding from lateral outline of head. Interfacetal setae long, dense. Tempus constricted towards base, about half as long as dorsal eye length. Head base rather narrow, subtruncate, slightly notched medially. Frontoclypeal suture or impression not observed. Head dorsal punctures rather small and shallow, denser on anterior half of head compared to vertex. Intervening spaces transversely microstrigose, as wide as to 3× as wide as punctures. Head dorsal setae yellowish, subdecumbent, moderately dense, directed posteriad. Tactile setae erect, long, present on head base and along lateral margin. Antenna slender, filiform, hardly extending towards base of pronotum when directed posteriad. Basal antennomere elongate, 1.5–1.6× as long as antennomere two. Antennomere three slightly longer than antennomere two, about 0.7× as long as antennomere four. Antennomeres 9–10 widened distally. Terminal antennomere long and slender, acutely pointed, about twice to 2.1× as long as penultimate antennomere. Terminal maxillary palpomere cultriform. Pronotum moderately glossy dorsally and on prothoracic hypomeron, flattened in dorsal aspect, elongate cylindrical, narrower than head across compound eyes. Anterior margin narrow, truncate. Lateral margins constricted laterally anteriad. Dorsal pronotal punctures distinctly larger, denser and deeper than those on head. Intervening spaces smooth to transversely microstrigose, generally as wide as to twice as wide punctures. Pronotal dorsal setation yellowish, subdecumbent, long, not concealing dorsal sculpture of pronotum. Tactile setae sparse, erect, no longer than ordinary setae. Scutellar shield small, truncate at posterior margin. Elytron elongate, moderately glossy, slightly and gradually narrows posteriad. Humerus broadly rounded. Postbasal transverse impression not indicated. Punctures on each elytron either confused or arranged into five to six incomplete, irregular rows; punctures distinctly smaller than those on pronotum. Intervening spaces about twice as wide as punctures, microstrigose. Elytral setation yellowish, moderately dense, long, directed posteriad, not concealing dorsal sculpture of elytron. Lateral humeral stria not shifted dorsad, not visible in dorsal view. Sutural stria complete, moderately broad. Metathoracic wing fully developed. Legs long. Tibial terminal spurs paired, serrate. Metatibial terminal spurs long. Tarsomeres of meso- and metathoracic leg elongated. Penultimate meso- and metathoracic tarsomere small. Basal metatarsomere longer than combined length of remaining tarsomeres. All claws strongly dentate at base. Male tergite VII broadly and shallowly emarginate at posterior margin ( Fig. 140 ). Male morphological sternite VII with subtruncate median protrusion at posterior margin ( Fig. 141 ). Tergite VIII, morphological sternites VIII and IX as in Fig. 142 . Aedeagus ( Figs 143–145 ) long and slender, basale about twice as long as apicale, paramere apically irregularly rounded, somewhat sinuous at external margin. Median lobe apically tripartite, its lateral prongs asymmetrical and longer than median prong.

Sexual dimorphism. Female ( Fig. 139 ) terminal antennomere twice as long as penultimate antennomere, compound eye somewhat comparatively shorter, frons 0.7–0.8× as wide as dorsal eye length, elytron not narrows posteriad. Female tergite VII broadly rounded at posterior margin ( Fig. 146 ), morphological sternite VII with rounded median protrusion at posterior margin ( Fig. 147 ).

Differential diagnosis. Due to the elongate, posteriad-narrowed head Macratria tongkonan sp. nov. is most similar to its Sulawesian congeners M. caerulescens Telnov, 2011 , M. maxbarclayi Telnov, 2011 and M. sanguiceps Telnov, 2011 but the dorsum without dark metallic blue reflection (present in M. caerulescens ), the intervening spaces on pronotum wider than punctures (narrower than punctures in M. maxbarclayi ), the longer tempus about half the eye length (the tempus about one fourth the eye length in M. sanguiceps ), and the terminalia and aedeagus different. Also appears externally similar to M. roosilehti sp. nov. (described above from the same locality) but differs in the comparatively stronger constricted, longer tempus, the comparatively narrower frons (cf. Fig. 78 & 138 ), the comparatively longer male terminal antennomere (about twice to 2.1× as long as penultimate antennomere in M. tongkonan sp. nov. versus about 1.2× as long in M. roosilehti sp. nov.), the comparatively less strongly punctured pronotal dorsum and elytra, the pale legs (the legs black to black-brown in M. roosilehti sp. nov.), and the different shape of the aedeagus.

Ecology. Collected from vegetation in disturbed lower montane rainforest.

Distribution. Central part of Sulawesi (administrative provinces of Central and South Sulawesi).

Figures

FIGURE 78. Macratria roosilehti sp. nov., holotype ♁, habitus, dorsal view.

FIGURES 138–139. Macratria tongkonan sp. nov. 138—Holotype ♁, habitus, dorsal view; 139—Paratype ♀, habitus, dorsal view [not to scale].

FIGURES 140–147. Macratria tongkonan sp. nov. 140—Holotype ♁, tergite VII, dorsal view; 141—Same, morphological sternite VII, ventral view; 142—Paratype ♁, morphological tergite VIII, morphological sternites VIII and IX, ventral view; 143—Same, aedeagus; 144—Holotype ♁, aedeagus, different position (less strongly pressed); 145—Paratype ♁, aedeagus, apical portion (strongly pressed); 146—Paratype ♀, tergite VII, dorsal view; 147—Same, morphological sternite VII, ventral view [not to scale].