Aulacopalpus aconcaguensis, Smith, 2002

Smith, Andrew B. T., 2002, Revision Of The Southern South American Endemic Genus Aulacopalpus Guérin-Méneville With Phylogenetic And Biogeographic Analyses Of The Subtribe Brachysternina (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini), The Coleopterists Bulletin 56 (3), pp. 379-437 : 401-403

Aulacopalpus aconcaguensis n. sp.

( Figs. 1 , 4 , 12 , Map 1)

Types. Holotype labeled a) ‘‘ CHILE: Aconcagua Cachagua 2­VIII­1974 P. Vidal & M. Donoso,’’ b) ‘‘ FMNH 986 L. Peña Coll. Acc #17–422.’’ Sixteen male paratypes with same data; one additional male paratype with identical locality label but without a second label. Two male paratypes labeled a) ‘‘HUAQUEN Costa Coqbo. 24, 31. Julio­1960 Coll: L. E. Pena,’’ b) ‘‘H. & A. HOWDEN COLLECTION ex. A. Martinez coll.’’ Two male paratypes labeled a) ‘‘TUNQUEN Costa Valpso 6, 15 Junio­1960 Coll: L. E. Pena,’’ b) ‘‘H. & A. HOWDEN COLLECTION ex. A. Martinez coll.’’ One male paratype labeled a) ‘‘ALGARROBO Costa Valpar. Julio­1957 Coll: L. E. Pena,’’ b) ‘‘H. & A. HOWDEN COLLECTION ex. A. Martinez coll.’’ One male paratype labeled a) ‘‘Q. AMOLANAS Costa Coqbo. 24, 31 Julio­1960 Coll: L. E. Pena,’’ b) ‘‘57,’’ c) ‘‘198,’’ d) ‘‘H. & A. HOWDEN COLLECTION ex. A. Martinez coll.’’ Two male paratypes labeled a) ‘‘ El Totoral Sa Antonio 28­V­ 1995 S. Rothmann,’’ b) ‘‘COLECCION SERGIO ROTHMANN.’’ One male paratype labeled ‘‘ CHILE SN. ANTONIO Las Cruces 20­VII­1976 col. G. Arriagada. ’’ One male paratype labeled a) ‘‘ CHILE SANTIAGO LAS CRU­ CES 20­VII­1976 Coll. G. ARRIAGADA,’’ b) ‘‘ CHILE SN. ANTONIO Las Cruces 20­VII­1976 col. G. Arriagada. ’’ One male paratype labeled a) ‘‘ CHILE V REGION Las Cruces Julio. 1998. leg. Gerardo Arriagada,’’ b) ‘‘COLEC­ CIÓN Gerardo Arriagada S. ’’ One male paratype labeled ‘‘ Fray Jorge Coquimbo 24­IX­1970 J. Solervicens. ’’ One male paratype labeled ‘‘ Co LA CAMPANA VALPARAISO 4–9–1979 J. Solervicens. ’’ One male paratype labeled ‘‘ Co Placenes Valpso. 1­VII­1968 Col. C. Vivar T. ’’ One male paratype labeled ‘‘P. PETORCA 2 km S. Zapallar 15­II­79 A. CHUBRETOVIC.’’ One male paratype labeled a) ‘‘FRAY JORGE Ovalle 27/VII/1987,’’ b) ‘‘ Leg. P. Vidal GH.’’ One male paratype labeled ‘‘182/1.’’

Holotype and eight male paratypes at FMNH. Seven male paratypes at MNNC. Six male paratypes at UNSM. Five male paratypes at CMNC. Three male paratypes at ABTS. Two male paratypes at PVGH. One male paratype each at BCRC, GASC, MLJC.

402 THE COLEOPTERISTS BULLETIN 56(3), 2002

Map 1. Distribution of Aulacopalpus aconcaguensis (•) and Aulacopalpus castaneus (O).

THE COLEOPTERISTS BULLETIN 56(3), 2002 403

Holotype. Male. Length 10.9 mm. Width 6.0 mm. Color: head and elytron light brown, pronotum dark brown with greenish reflections. Head: Figure 4 . Dorsal surface densely

punctate and moderately setose; punctures moderately­sized; setae long, slender, tawny. Frontoclypeal suture weakly bisinuate. Labrum with apex vertically produced with respect to clypeus, weakly produced at middle, rounded. Maxillary palpus with terminal segment enlarged, with deep sulcus. Mentum with surface moderately setose, apex acuminate. Antenna 10­segmented (segments 5 and 6 partially fused making antenna appear 9­segmented); club elongated, flattened, longer that combined basal segments (club length approximate to head length). Pronotum: Form widest at middle, midline absent (weakly punctate in other individuals). Surface densely punctate, moderately setose; punctures moderately­sized; setae long, slender, tawny. Scutellum: Shape parabolic: 1.3 times wider than long medially. Surface densely punctate, setose; punctures moderately large; setae long, slender, tawny. Elytron: Surface glabrous. Longitudinal striae punctate; punctures moderately­sized, separated by 2–6 puncture widths. Intervals sparsely punctate with moderately­sized punctures. Epipleuron with row of dense setae ventral to bead. Pygidium: Width 2.3 times length medially. Surface sparsely punctate, moderately setose; punctures moderately­sized; setae tawny. Venter: Thorax densely setose; setae long, tawny. Legs: Protibia with 3 subequally large teeth. Tarsal claws with modified claw slightly thickened when compared to other claw, apex bifurcate (similar to Fig. 7 ). Tarsomere 5 without ventromedial tooth (similar to Fig. 6 ). Meso­ and metatibia with apical spurs slender, acute. Mesotibial apex with 17 spinules. Metatibial apex with 25 spinules. Parameres: Figure 12 . Basally with broad, medial furrow.

Variation. (34 paratypes). Length 10.9–16.6 mm. Width 6.0– 8.6 mm. Color of elytron light to dark brown. The paratypes do not differ significantly from the holotype. Female unknown.

Etymology. From the latinized adjective for ‘‘Aconcaguan,’’ referring to the region in Chile in which this species is found and the province where most of the type series was collected.

Diagnosis. This species is distinguished from all other species in the genus Aulacopalpus by the following combination of characters: antennal club elongated in males (length equal to or slightly shorter than head); pronotum green to greenish brown when viewed without magnification; elytron brown, glabrous; all tarsal claws with modified claw slightly thickened when compared to other claw and with apex bifurcate (males); tarsomere 5 without ventromedial tooth; parameres basally with broad, medial furrow.

Distribution (Map 1). Coastal Chile from Coquimbo to Santiago.

Locality Data. Based on 35 specimens examined from the CMNC, FMNH, MNNC, PVGH .

CHILE (34). ACONCAGUA (19): Cachagua, Zapallar . COQUIMBO (5): Coquimbo , Guaquén, Ovalle, Quebrada Amolanas . SANTIAGO (3): Las Cruces . VALPARAÍSO (7): Algarrobo , El Totoral, La Campana, Placeres Tunquén .

NO DATA (1).

Temporal Data. February (1), May (2), June (2), July (9), August (18), September (2).

Figures

Fig. 1. The Brachysternina. First row: Aulacopalpus aconcaguensis (male paratype), A. aconcaguensis (male paratype), A. castaneus (male), A. castaneus (male), A. castaneus (male), A. punctatus (male). Second row: A. ciliatus (male), A. clypealis (male),

Figs. 4–5. 4) Head of male Aulacopalpus aconcaguensis. 5) Head of male Aulacopalpus viridis.

Figs. 12–20. Parameres in caudal view. 12) Aulacopalpus aconcaguensis; 13) Aulacopalpus ciliatus; 14) Aulacopalpus clypealis; 15) Aulacopalpus pilicollis; 16) Aulacopalpus castaneus; 17) Aulacopalpus punctatus; 18) Aulacopalpus viridis; 19) Aulacopalpus pygidialis, 20) Aulacopalpus valdiviensis.

Figs. 6–11. 6) Protarsomere 5 and claw of male Aulacopalpus pilicollis, 7) Protarsomere 5 and claw of male Aulacopalpus pygidialis. 8) Protarsomere 5 and claw of female Aulacopalpus ciliatus. 9) Dorsal view of modified proclaw of male Aulacopalpus ciliatus. 10) Dorsal view of modified proclaw of male Aulacopalpus castaneus. 11) Lateral view of Aulacopalpus valdiviensis parameres and phallobase.