Crossopriza kandahar, Huber, 2022

Huber, Bernhard A., 2022, Revisions of Holocnemus and Crossopriza: the spotted-leg clade of Smeringopinae (Araneae, Pholcidae), European Journal of Taxonomy 795 (1), pp. 1-241 : 168-172

publication ID 10.5852/ejt.2022.795.1663

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scientific name

Crossopriza kandahar

sp. nov.

Crossopriza kandahar View in CoL sp. nov.

Figs 353A View Fig , 664–680 View Figs 664–666 View Figs 667–675 View Figs 676–680


Males are easily distinguished from known congeners by presence of small pair of apophyses frontally on male chelicerae ( Fig. 669 View Figs 667–675 ); also by details of male palp ( Figs 668, 671 View Figs 667–675 ; procursus tip with small dorsal flat process; distal bulbal sclerite with two large prolateral apophyses), and by shape of epigynum ( Fig. 677 View Figs 676–680 ; short and wide plate, pair of large pockets close together but not on median rim; similar to C. sengleti sp. nov.).


The species name refers to the type locality; noun in apposition.

Type material

Holotype AFGHANISTAN • ♂; Kandahar Province, W of Kandahar; 31.62° N, 65.60° E; 1000 m a.s.l.; 31 Jul. 1975; A. Senglet leg.; MHNG. GoogleMaps

Other material examined

AFGHANISTAN • 10 ♂♂, 15 ♀♀, 4 juvs (2 vials); same collection data as for holotype; MHNG GoogleMaps .


Male (holotype)

MEASUREMENTS. Total length 5.3, carapace width 1.65. Distance PME–PME 70 µm; diameter PME 110 × 150 µm; distance PME–ALE 30 µm; diameter AME 95 µm; distance AME–AME 30 µm. Leg 1: 37.5 (10.8 + 0.7 + 10.3 + 13.3 + 2.4), tibia 2: 7.2, tibia 3: 5.6, tibia 4: 6.8; tibia 1 L/d: 61; femora 1–4 diameters: 0.26, 0.23, 0.22, 0.24.

COLOR (in ethanol). Carapace ochre-yellow, anteriorly in median pit light brown; sternum with many small brown spots partly arranged in radial marks; legs ochre-yellow, without darker rings, with small black lines on femora and (few) on tibiae; abdomen pale gray, with whitish internal marks and indistinct dark band posteriorly above spinnerets; ventrally with broken dark band, with three indistinct parallel longitudinal marks behind gonopore.

BODY. Habitus similar to C. tiwi sp. nov. (cf. Figs 393–394 View Figs 391–398 ). Ocular area slightly raised. Deep thoracic pit and pair of furrows diverging from pit toward posterior margin. Clypeus unmodified, only rim slightly more sclerotized than in female. Sternum wider than long (1.1/0.75), unmodified. Abdomen slightly elongated, dorso-posteriorly angular to conical.

CHELICERAE. As in Figs 669–670 View Figs 667–675 , with two pairs of frontal apophyses; proximal pair small and simple; distal pair larger, at tip with one large modified cone-shaped hair each; distance between tips of modified hairs 180 µm; lateral stridulatory ridges clearly visible in dissecting microscope.

PALPS. As in Figs 664–666 View Figs 664–666 ; coxa with rounded retrolateral hump; trochanter barely modified; femur distally strongly widened, with rounded ventral protrusion, proximally with prolateral stridulatory pick, without retrolateral transversal line, without retrolateral proximal process; femur-patella joints shifted toward prolateral side; tibia-tarsus joints slightly shifted toward retrolateral side; tarsus without macrotrichia; procursus ( Figs 667–668 View Figs 667–675 ) straight, dorsal hairs not curved, prolateral proximal hump set with many hairs, procursus tip with strong ventral sclerite, small dorsal sclerite with prolateral-dorsal ridge, and membranous elements; genital bulb ( Figs 671–673 View Figs 667–675 ) with simple basal sclerite connected to distal (main) sclerite, sperm duct opening not seen; distal sclerite with retrolateral ridge and distinctive set of two prolateral apophyses and ridge.

LEGS. Femur 1 with single row of ~18 ventral spines; without curved hairs; few vertical hairs; retrolateral trichobothrium of tibia 1 at 3.5%; prolateral trichobothrium absent on tibia 1, present on other leg tibiae; tarsal pseudosegments not seen.

Male (variation)

Tibia 1 in 11 males (incl. holotype): 9.6–10.8 (mean 10.0). Some males with few indistinct dark marks dorsally on abdomen; ventral abdominal marks behind gonopore variable (2–4 parallel bands).


In general similar to male but without spines on legs, apparently without stridulatory files on chelicerae, and with stridulatory organ consisting of pair of weakly sclerotized but distinct processes posteriorly on carapace and pair of distinct light brown plates anteriorly on abdomen. Tibia 1 in 15 females: 8.1–10.0 (mean 9.1). Epigynum as in Figs 676–677 View Figs 676–680 , main epigynal plate triangular, short but wide, weakly protruding, only posteriorly heavily sclerotized; with pair of large pockets 80 µm apart; internal sclerotized arcs and median anterior internal sclerite visible in uncleared specimens; posterior plate light brown, short but wide. Internal genitalia ( Figs 674–675 View Figs 667–675 , 678–680 View Figs 676–680 ) with large oval pore plates converging anteriorly, pores arranged in groups, dorsal arc strong but simple, ventral arc with median sclerite (rather than pouch or pocket) of unknown function.


Known from type locality only, in Afghanistan, Kandahar Province ( Fig. 353A View Fig ).


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