Gossia pancheri (Brongn. & Gris) N. Snow

Snow, Neil, 2020, A revision of New Caledonian Gossia N. Snow & Guymer (Myrtaceae), Adansonia 42 (7), pp. 131-177 : 166-168

Gossia pancheri (Brongn. & Gris) N. Snow ( Figs 16D ; 20 ; 21A-D )

Austrobaileya 8: 181 (2010). — Eugenia pancheri Brongn. & Gris, Bulletin de la Société botanique de France 12: 180 (1865). — Austromyrtus pancheri (Brongn. & Gris) Burret, Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem 15: 503 (1941).

— Typus: New Caledonia. Grande Terre, South Prov ., Montagnes de Yaté , Vieillard 508 (lecto- [designated by Snow & Veldkamp 2010], P [ P00462968 ]; isolecto-, P[P00462969, P00469970, P00602555]) . — Snow annotated the lectotype in 2004 but did not specify a barcode number ( Snow & Veldkamp 2010: 181), which at that time did not exist. One of the isolectotypes (P00602555) was annotated by the author in 2005 mistakenly as “lectotype”, instead (correctly) of “isolectotype”.

Myrtus luteo-viridis Baker f., The Journal of the Linnean Society, Botany 45: 312 (1921). — Austromyrtus luteo-viridis (Baker f.) Burret, Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem 15: 505 (1941). — Typus: New Caledonia. Grande Terre, South Prov ., Ngoye Mts. to NW, Compton 1339 (holo-, BM [ BM000581650 ], photo at BISH]).

Eugenia angustibracteolata Baker f., The Journal of the Linnean Society, Botany 45: 313 (1921). — Typus: New Caledonia. Grande Terre, South Prov ., Presqu’île Bogota , 13.V.1914, Compton 1002 (holo-, BM [ BM000581654], photo at BISH).

SPECIMENS ECAMINED. — New Caledonia. North Prov., Boulinda près campement, 500 m, 23.IV.1968, Jaffré 18 (NOU, P); Massif du Ko-

niambo, 250 m, 18.XI.1971, Jaffré 516 (NOU[NOU029667]); Massif du Koniambo, 900 m, 20.XII.1972, Jaffré 1038 ( NOU [ NOU 29662]) ; Kouaoua : Mea, 12.I.1978, Jaffré 2275 ( NOU [ NOU029649 ]) ; Mont Boulinda , pente au-dessus de la Oua-Népoua, 750-850 m, 26.VII.1967, MacKee 17177 ( NOU [ NOU029683 ]) ; Taom , Mt. Homédéboa, 800- 900 m, X.1969, MacKee 20995 ( NOU [ NOU029680 ]) ; Mont Do , 950-1020 m, X.1969, MacKee 21089 ( NOU [ NOU029677 ) ; Manambo , 500 m, 29.IX.1989, MacKee 44623 ( NOU [ NOU0029652 ], P[P02089832]) ; Poro , Kaéoua, 500 m, 17.XII.1992, MacKee 46074 (P[P02089865]) ; Contrefort Nord-Ouest du Mt. Taom, 5.X.1979, MacKee 36887 (P); Plateau de Ouazangou , 26.III.1980, MacKee 37951 (P[P02089833]) ; Mont Taom (Crête Ouest), 900 m, 8.I.1981, MacKee 38519 ( NOU [ NOU029650 ], P[P02089884]) ; Mont Taom , 900 m, 31.VII.1981, MacKee 39418 (MO, NOU [ NOU029647 ], P[P02089883]) ; Poro , Kaseoua, 500 m, 17.XII.1992, MacKee 46074 ( NOU [ NOU029653 ], P[P02089865]) ; Mt Koniambo , c. 700 m, 13.X.1982, McPherson 5019 (MO, NOU [ NOU053242 ]) ; Mt Koniambo , c. 550 m, 5.I.1983, McPherson 5297 ( BRI, MO, NOU [- NOU029654 ], NSW, PTBG, WELTU) ; Mt. Colnett , 800-925 m, 2.XI.2003, McPherson 19142 ( BISH, MO, NOU [ NOU004366 ], P) ; Est du Kopeto, 900 m, 28.XI.1980, Morat 6642 ( NOU [ NOU029671 ]) ; Massif du Tchingou, face est, 950 m, 29.III.2001, Munzinger 563 (MO, NOU [ NOU029668 ]) ; Massif du Tchingou, face est, 1000 m, 31.III.2001, Munzinger 588 (MO); Massif du Tchingou, face est, 1120 m, 31.III.2001, Munzinger 615 (MO, P) ; Paéoua Mt , 800-1100 m, 26.X.2010, Munzinger 6017 (MO, P). — South Prov., Pente brûlée près de la mine du “Mois de Mai”, 300 m, 13.VIII.1951, Baumann- Bodenheim 15115 (A, US, Z) ; Canala , Presqu’île de Bogota, c. 450 m, 3.VII.1997, Dagostini 120 ( NOU [ NOU029675 ]) ; Goro-Nickel , relevé 34, 20. VI.2002, Dagostini & Rigault 505 ( NOU [ NOU029679 ]) ; Goro , concession SLN “Camille”, 18. V.2002, Dagostini 615 (MO, NOU [ NOU029678 ]) ; Col de Mouirange , forêt Desmazure, 10.III.2005, Dagostini 1050 ( NOU [ NOU009280 ]) ; Montagne des Sources , 3.XII.1992, Dawson WELTU 16573 ( NOU [ NOU029676 ], WELTU) ; Bord de la Fausse Yaté, 10.VII.1981, Hoff 3581 ( NOU [ NOU029659 ]) ; Dans la partie supérieure de la vallée de l’Odjijoni, 180 m, 2. VI.1951, Hürlimann 1463 ( US, Z) ; Vallée latérale de la Pourina, 160 m, 4. VI.1951, Hürlimann 1482 ( US, Z) ; au-dessus du campement Bernier (Montagne des Sources), 850 m, 13.XI.1951, Hürlimann 3019 ( US, Z) ; Plaine des Lacs , 15.IV.1969, Jaffré 18 ( NOU [ NOU029660 ]) ; Plaine des Lacs , 15.IV.1969, Jaffré 200 ( NOU [ NOU029658 ]) ; Région de Kouaoua , 30.IX.1969, Jaffré 275 ( NOU [ NOU 29661]) ; Kouaoua , 30.IX.1969, Jaffré 294 ( NOU [ NOU029664 ]) ; Mt. Kouakoué , 21°58’43”S, 166°32’15”E, 980-1050 m, 9.X.2004, Labat 3486 (K n.v., MO, NOU [ NOU009917 ], P[P00454747], WELTU) ; 12 km S of Riv. des Pirogues, ridge overlooking Prony , 200 m, 15.X.1955, MacKee 3247 ( US) ; Yaté road near barrage, 150-200 m, 6. V.1956, MacKee 4494 (L) ; Route de la Montagne des Sources , 500-700 m, 27.X.1956, MacKee 5665 (L) ; Vallée de Mamié , 200 m, 6.X.1966, MacKee 15740 ( NOU [ NOU029681 ]) ; Ouroué , 21.VII.1967, MacKee 17148 (P[P02089831]) ; Rivière du Humboldt, 800-900 m, 29.X.1967, MacKee 17838 ( NOU [ NOU029648 ]) ; Thio-Mt Nékandi , 1200- 1300 m, 7.XI.1967, MacKee 17925 ( NOU [ NOUO29862 ]) ; Mé Ori , Plateau sommital, 900-1000 m, 3.XII.1969, MacKee 21205 ( NOU [ NOU029657 ]) ; Vallée de la Ouinnée, 25. V.1974, MacKee 28691 ( NOU [ NOU029666 ]) ; Yaté , plateau au Sud du village, 300 m, 4. VI.1977, MacKee 33235 ( NOU [ NOU029665 ]) ; Dalmates Les (Mouirange Col), 150 m, 29.II.1980, MacKee 37865 ( NOU) [ NOU029608 ] ); Vallée de la Ni, 100 m, 3.XII.1981, MacKee 40035 ( NOU [ NOU029651 ]) ; Hauteurs de Yaté , 250 m, 10.III.1985, MacKee 42493 (MO, NOU [ NOU029669 ], P[P02089882]) ; Along old lumber road to top of Mt Mé Ori, above Katrikoin, 8.IX.1980, McPherson 3069 ( BRI, MO, NOU [ NOU029646 ], NSW, WELTU) ; Montagne des Sources , c. 700 m, 25.X.1980, McPherson 3275 ( BRI, MO, NOU [ NOU029663 ], NSW, WELTU) ; Along Nouméa-Yaté road, near descent towards Yaté, 400 m, 8.VII.1981, McPherson 3894 (MO, NOU [ NOU 20670], WELTU) ; Upper reaches of valley Rivière des Pirogues, N of Nouméa-Yaté road, c. 300 m, 12.XII.1981, McPherson 4451 (MO, NOU [ NOU029656 ], NSW, WELTU) ; by Fausse Yaté River , 23.XI.1982, McPherson 5185 ( BISH, MO) ; Forested slopes above a tributary of the Rivière Ni, 750-770 m, 12.XI.2003, McPherson 19230 ( BISH, MO, NOU) ; Me Adeo piste en direction du Menazi, 24.III.1988, Morat 8008 ( NOU [ NOU029655 ]) ; Sommet du Col de Yaté , 26.VII.1965, Schmid 506 ( NOU [ NOU029672 ]) ; Mine Pauline , vers 450 m, 18.III.1968, Veillon 1648 ( NOU [ NOU029673 ]) ; Route vers Prony, après Col de Crève-Coeur, 14.X.1968, Veillon 1859 ( NOU [ NOU029674 ], P[P00462967, P05094569], WELTU). — Ab loco. Deplanche 530, 1861 (P) ; Deplanche 531, 1861 (P); Deplanche 64172, 1861 (P).

REPRESENTATIVE HERBARIUM SPECIMEN. — P02089884.

ETYMOLOGY. — Named for Jean Armand Isidore Pancher (1814‒1877), French botanist who worked in New Caledonia.

DISTRIBUTION, HABITAT AND PHENOLOGY. — Widespread on Grande Terre ( Fig. 20 ); in maquis, forsest edges, and humid forests, often on ultramafics, 150-1300 m. Flowering throughout the year; fruiting mostly December through July.

DESCRIPTION

Shrubs or trees

1.5- 10 m.

Branchlets

Terete to compressed, surface developing small dark rectangular flakes; internodes> 5 mm.

Leaves

Coriaceous, 2 per node; petioles 2-6.5 mm, sulcate; blades 4.0-8.5 × 1.7-3.8 cm, elliptic, base cuneate, surface flat, margin slightly revolute, apex obtuse or retuse, midnerve above sulcate, secondary veins above raised prominently, oil glands of lower surface dense (and of uniform size), prominent.

Inflorescence

2-5.5(-8) cm, pedicels (1.5-) 5-9 mm, a simple cyme, raceme, or panicle, terminal or axillary; extrafloral bracts leafy.

Bracteoles

1.3-2 mm, linear to narrowly elliptic, sparsely to densely sericeous.

Hypanthium

c. 3 mm × 3-3.5 mm, campanulate, surface smooth, sericeous to densely sericeous; ovary apex densely tomentose.

Calyx

Lobes 5, 1.3-2.5 mm, broadly rounded to almost truncate, sericeous to densely sericeous above and below, green.

Petals 4.5-6 × c. 5 mm, somewhat tomentose above and below.

Filaments 2.5-4 mm; anther sacs 0.5-0.8 mm.

Style

c. 4 mm, lower half villous. Fruits (material scant) 5-6 × 5-6 mm, dark blue-black, globose, base rounded; seeds 1-2.

REMARKS

Gossia pancheri is the second most common member of the genus in New Caledonia after Gossia vieillardii . It is common in the southern part of Grande Terre but ranges north to Mt. Colnett.

The best distinguishing characters include the raised secondary and tertiary venation of the leaves, the thickly coriaceous texture and often glossy sheen of the upper leaf surface.

The species most closely resembles Gossia kaalaensis sp. nov., but that species has narrower leaves, a more acute leaf apex, and a glabrous ovary apex. Some specimens (e.g., MacKee 28691 and 42493) have more densely pubescent and somewhat smaller flowers, but these differences are merely quantitative.

Figures

FIG. 16. — A-C, Gossia kuakuensis (Baker f.) N. Snow vel aff.: A, inflorescences and leaves;B, leaves and branchlets with dark rectangular flaking bark; C, young fruits; D, Gossia pancheri. Vouchers: A-C, Fleurot 226 (NOU, P); photo ©Dominique Fleurot, with permission; from Näräjaa, Ponérihouen (http://endemia.nc); D, J.-N. Labat 3486 (P).

FIG. 20. — Distribution maps of Gossia pancheri (Brongn. & Gris) N. Snow, G. ramiflora N. Snow, sp. nov. and G. virotii (Guillaumin) N. Snow.

FIG. 21. — A-D, Gossia pancheri (Brongn. & Gris) N. Snow: A, flowering branch with details of abaxial leaf venation (offset on right); B, detail of flower prior to anthesis; C, longitudinal section of flower bud showing bilocular ovary with axile placentation; D, leaf and fruits; E-I, Gossia ramiflora N. Snow, sp. nov.: E, flowering branch; F, details of more or less rectangular flaking bark; G, flowers prior to anthesis; H, detail of flower bud; I, another view of leaf (with detail of sinuous margin) with young fruits. Vouchers: A-C, McPherson 3275 (WELTU); D, McPherson 5019 (NOU); E-H, MacKee 12272 (WELTU); I, Phillips 2105 (NOU). Scale bars: A, D, I, 2 cm; B, C, H, 3 mm; E, 2 mm; F, G, 5 mm.