Acomoptera echinosa, Kerr, Peter H., 2011

Kerr, Peter H., 2011, Six new species of Acomoptera from North America (Diptera, Mycetophilidae), ZooKeys 137, pp. 41-76 : 51-54

Acomoptera echinosa sp. n. Figs 1214

Type Material.

Holotype: ♂, "USA: OR: Lincoln Co., Waldport, Malaise trap, 44.4266°N, - 124.0513°W, 56masl, 1-15.ix.2010, John D. Pinto CSCA11L044" / "HOLOTYPE 11G099 ♂ Acomoptera echinosa Kerr 2011" [red label]. Deposited in CSCA, complete specimen in excellent condition, mounted on gray point.

Paratypes: 1 ♂, "USA: CA: Humboldt Co., Patrick's Point SP, redwood grove behind visitor center, 41°08.11'N, 124°09.28'W, ~10masl, 10.iv.2008-12.ii.2009 P. Kerr, P. Haggard CSCA09L117" [CSCA] 1 ♂, "Canada: B.C., Upper Carmanah Valley, 4. vii– 15.vii.1991 N. Winchester, FF.MT4" [48.67°, -124.69°; CASC] 1 ♂, "Canada: B.C., Upper Carmanah Valley, 28. viii– 9.ix.1991 N. Winchester, FF.MT1" [48.67°, -124.69°; CASC].

Diagnosis.

This species is similar to Acomoptera nelsoni sp. n. in that the male gonocoxites display a prominent dorsomedial comb (Fig. 14A) and the outer surface of the gonostyli bear denticulations (Figs 14B, 14D). The gonostyli of both species are also similar in that the dorsoapical margin comes to an acute point and is darker in color than the rest of the gonostylus (Fig. 14D). The gonostyli of Acomoptera echinosa sp. n., however, lack denticulations on the inner surface of the gonostyli dorsally (Figs 13A, 14A).

Description.

Male. Body length (n=2): 6.5-6.9 mm (avg = 6.7 mm). Wing length: 4.8-5.7 mm (avg = 5.3 mm).

Coloration (Fig. 12). Head brown; palpomeres light brown. Antennal scape light brown, pedicel yellowish, base of first flagellomere yellowish, otherwise flagellomeres brown. Thorax cream-colored to brown; scutum brown to dark brown; dorsocentral areas of scutum defined by lighter brown coloration, scutum setae gold- or golden brown-colored; laterotergite and mediotergite light brown to cream-colored. Legs becoming increasingly brown towards tarsi, coxae cream-colored; femur yellowish or cream-colored, tibia yellowish brown to brown, tarsi brown. Wing hyaline without markings, wing veins light brown; haltere stem and knob cream-colored to light brown. Abdominal segments concolorous, brown, slightly darker posteriorly, with golden brown setae. Terminalia yellowish brown to brown.

Head. Ocelli slightly raised; middle ocellus clearly smaller than (approx. 0.25 × size of) lateral ocelli, lateral ocellus located approx. width of ocellus or less from eye margin, separated from median ocellus by approx. twice its own diameter or a little more. Eyes with sparse, inconspicuous microsetae, which are approximately as long as width of facet. Face with golden brown setae, longest of which approx. same length as width of face. Antenna and abdomen subequal in length. Palpus approx. 1 × width of head (anterior view); length of palpomeres 2 and 3 nearly subequal (palpomere 3 longer); palpomere 4 approx. 6 × longer than wide; palpomere 5 approx. 11 × longer than wide, subequal to or shorter than combined length of palpomeres 3 and 4.

Thorax. Antepronotum bearing setae; remaining thoracic sclerites bare. Tarsal claw usually with one small ventral tooth. Wing venation similar to others in the genus (e.g., Acomoptera digitata sp. n., Fig. 8); costal vein extends beyond R5, approx. 0.33 × distance between R5 and M1; R1, R5, and M1 with at least some setae on lower surface (ventral M2 sometimes with setae, also).

Male Genitalia (Figs 13-14). Epandrium approx. 2 × wider than long (Fig. 14F). Gonocoxites with developed dorsomedial comb (Figs 14A, 14C). Gonostylus complex, as shown in Figs 13, 14 A–B, 14 D–E.

Female unknown.

Etymology.

The species epithet "echinosa" is an adjective derived from the Greek word meaning hedgehog or sea-urchin, referring to the spiny surface of the gonostylus.