Clitobius Mulsant & Rey, 1859

Purchart, Luboš & Kamiński, Marcin Jan, 2017, A taxonomic review of the genus Clitobius with description of a new species from Oman (Coleoptera: Tenebrionidae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57, pp. 139-163 : 142-143

Genus Clitobius Mulsant & Rey, 1859

( Figs 1 , 2 , 3C–I, 4)

Clitobius Mulsant & Rey, 1859: 141 (original description). Type species: Clitobius sabulicola Mulsant & Rey, 1859

(= Opatrum ovatum Erichson, 1843 ). MULSANT & REY (1860): 49 (repeated description); FAIRMAIRE (1885): ccvi

(catalogue); SEIDLITZ (1891): 130 (key to Tenebrionidae of Transylvania); SEIDLITZ (1898): 410 (key to Opatrini ), 452 (key to Melanimonina ); CHAMPION (1895): 111 (catalogue); REITTER (1904): 136 (key to Opatrini ), 178

(new subgenus); GEBIEN (1910): 340 (catalogue); REICHARDT (1936): 81 (key to Palaearctic Opatrini ), 200 (key to subgenera), 201 (key to species); ESPAÑOL (1943): 142 (key to Clitobius subgenera); ESPAÑOL (1944): 30 (key to palaearctic Opatrina ) ; ESPAÑOL (1962a): 219 (key to Opatrinae of Canary Islands) ; HAR. LINDBERG (1950): 12

(new species); KOCH (1959a): 11 (definition and key of Clitobius group of genera); KOCH (1959b): 592 (key to

Clitobius group of genera); KOCH (1960): 391 (key to Apteroclitobius species ), 403 (key to Clitobius generic group); ABDURAKHMANOV & NABOZHENKO (2011): 103 (key to Opatrina).

Apithesis Waterhouse, 1881: 476 (syn. nov.). Type species: Apithesis obesa Waterhouse, 1881 . GAHAN (1903): 280

(repeated description; with figure of habitus); GEBIEN (1940): 781 [588] (catalogue); KOCH (1970): 115; SCHA-

WALLER (2004): 455 (new records); LÖBL et al. (2008): 302 (catalogue). Apteroclitobius Koch, 1960: 391 (syn. nov.). Type species: Clitobius grandis Fairmaire, 1896 . Halonomus Wollaston, 1861: 201 (syn. by FAIRMAIRE 1885: ccv). Type species: Halonomus grayii Wollaston, 1861 .

WOLLASTON (1864): 490; WOLLASTON (1865): 415 (discussion); WOLLASTON (1867): 191 (redescription); ALLARD

(1883): 30 (key to species); IWAN & LÖBL (2007): 733 (type species designation). Halanomus [sic!]: ESCALERA (1914): 339. Note. The above-listed publications concern only the most important nomenclatural acts and taxonomic works. For justification of the newly proposed synonymies see the discussion.

Diagnostic characters. According to KOCH (1960) and subsequent authors (e.g. SCHAWALLER 2009) Clitobius together with Ammidium Erichson, 1843 , Diaderma Koch, 1960 , Falsammidium Koch, 1960 , Falsocaedius Español, 1943 , and Freyula Koch, 1959 form a separate generic complex. All these taxa, together with some other genera representing Opatrini , were recently classified within the restituted subtribe Ammobiina (see IWAN & KAMIŃSKI 2016). Taxonomic distinctiveness of Clitobius within the above-mentioned generic complex is grounded in the following characters: base of pronotal disc bisinuate, with complete margination ( Figs 2B vs 3A); prosternal apophysis, in lateral view, horizontal, not gradually depressed or saddle-like (Figs 3B vs 3C); sides of pronotum and elytra glabrous or covered with fine and short setae; primary rows coarse ( Figs 2F vs 3A); protibia with relatively small apical tooth and without median tooth ( Fig. 1 ).

Figures

Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view).

Fig. 2. Detailed morphology of Clitobius obesus (Waterhouse, 1881) (A–C, F) and C. omanicus sp. nov. (D, E, G). Abbreviations: a – trochanter base; b – femur base; c – rounded posterior angle of pronotal disc; d – not depressed sides of pronotal disc; e – convex intervals; f – ventral; g – dorsal; h – lateral view.